[1] HUNTON P.Research on eggshell structure and quality:an historical overview[J].Revista Brasileira de Ciência Avícola,2005,7(2):67-71.

[2] NARUSHIN V G,ROMANOV M N.Egg physical characteristics and hatchability[J].World's Poultry Science Journal,2002,58(3):297-303.

[3] ROLAND D A.Research note:egg shell problems:estimates of incidence and economic impact[J].Poultry Science,1988,67(12):1801-1803.

[4] MABE I,RAPP C,BAIN M M,et al.Supplementation of a corn-soybean meal diet with manganese,copper,and zinc from organic or inorganic sources improves eggshell quality in aged laying hens[J].Poultry Science,2003,82(12):1903-1913.

[5] BAR A,VAX E,STRIEM S.Relationships among age,eggshell thickness and vitamin D metabolism and its expression in the laying hen[J].Comparative Biochemistry and Physiology Part A:Molecular & Integrative Physiology,1999,123(2):147-154.

[6] NYS Y.Recent developments in layer nutrition for optimizing shell quality[C]//Proceedings of the 13th European Symposium on Poultry Nutrition.Blankenberge,Belgium:ESPN,2001:45-52.
[7] 张亚男,武书庚,张海军,等.蛋壳品质营养调控的研究进展[J].中国畜牧杂志,2012,48(21):79-83.
[8] 张亚男,王晶,武书庚,等.糖胺聚糖对蛋壳品质的调控[J].动物营养学报,2015,27(11):3332-3337.
[9] 张亚男,齐晓龙,武书庚,等.锰、锌在蛋壳品质调控中的应用[J].饲料工业,2013(增刊):60-63.
[10] YANG X J,ZHONG L L,AN X F,et al.Effects of diets supplemented with zinc and manganese on performance and related parameters in laying hens[J].Animal Science Journal,2012,83(6):474-481.

[11] CORNESCU G M,CRISTE R D,UNTEA A E,et al.Supplementation of manganese and zinc in laying hens diet improves eggshell quality[J].Lucrãri Ştiintifice-Seria Zootehnie,2013,60:29-34.
[12] 张亚男,武书庚,张海军,等.锌添加水平对蛋鸡生产性能和蛋壳品质的影响[J].动物营养学报,2013,25(5):1093-1098.
[13] 张亚男,齐晓龙,武书庚,等.硫酸锌和蛋氨酸锌对产蛋后期蛋鸡生产性能、蛋品质及抗氧化性能的影响[J].动物营养学报,2013,25(12):2873-2882.
[14] 张亚男.饲粮锌对产蛋后期蛋鸡蛋壳品质及抗氧化机能的影响[D].硕士学位论文.北京:中国农业科学院,2013.
[15] NIELSEN F H.History of zinc in agriculture[J].Advances in Nutrition:an International Review Journal,2012,3(6):783-789.

[16] Panel on Additives and Products or Substances Used in Animal Feed (FEEDAP).Scientific opinion on the potential reduction of the currently authorised maximum zinc content in complete feed[J].EFSA Journal,2014,12(5):3668.
[17] ZAMANI A,RAHMANI H R,POURREZA J.Supplementation of a corn-soybean meal diet with manganese and zinc improves eggshell quality in laying hens[J].Pakistan Journal of Biological Sciences,2005,8(9):1311-1317.

[18] NETO M A T,PACHECO B H C,ALBUQUERQUE R,et al.Dietary effects of chelated zinc supplementation and lysine levels in ISA Brown laying hens on early and late performance,and egg quality[J].Poultry Science,2011,90(12):2837-2844.

[19] SAZZAD H M,BERTECHINI A G,NOBRE P T C.Egg production,tissue deposition and mineral metabolism in two strains of commercial layers with various levels of manganese in diets[J].Animal Feed Science and Technology,1994,46(3/4):271-275.
[20] FASSANI Š J,BERTECHINI A G,DE OLIVEIRA B L,et al.Manganese in nutrition of leghorn hens in the second cycle of production[J].Ciência e Agrotecnologia,2000,24(2):468-478.

[21] XIAO J F,ZHANG Y N,WU S G,et al.Manganese supplementation enhances the synthesis of glycosaminoglycan in eggshell membrane:a strategy to improve eggshell quality in laying hens[J].Poultry Science,2014,93(2):380-388.

[22] HOSSAIN S M,BERTECHINI A G.Effect of varying manganese and available phosphorus levels in the diet on egg production and eggshell quality of layers[J].Animal Feed Science and Technology,1998,71(3/4):303-310.
[23] OUZZINE M,GULBERTI S,NETTER P,et al.Structure/function of the human Galβ1,3-glucuronosyltransferase:dimerization and functional activity are mediated by two crucial cysteine residues[J].The Journal of Biological Chemistry,2000,275(36):28254-28260.
[24] 肖俊峰.日粮锰源和水平对产蛋鸡蛋壳品质的影响及其机理[D].博士学位论文.北京:中国农业科学院,2014.
[25] LEACH R M,Jr.,GROSS J R.The effect of manganese deficiency upon the ultrastructure of the eggshell[J].Poultry Science,1983,62(3):499-504.

[26] XIAO J F,WU S G,ZHANG H J,et al.Bioefficacy comparison of organic manganese with inorganic manganese for eggshell quality in Hy-Line Brown laying hens[J].Poultry Science,2015,94(8):1871-1878.

[27] CHOWDHURY S D.Shell membrane protein system in relation to lathyrogen toxicity and copper deficiency[J].World's Poultry Science Journal,1990,46(2):153-169.

[28] BAUMGARTNER S,BROWN D J,SALEVSKY E,et al.Copper deficiency in the laying hen[J].The Journal of Nutrition,1978,108(5):804-811.
[29] 武书庚.日粮中不同硫酸铜和柠檬酸铜添加水平及其组合对产蛋鸡生产性能及蛋品质的影响[D].硕士学位论文.北京:中国农业科学院,2001.
[30] PEKEL A Y,DEMIREL G,ALP M,et al.Influence of different dietary copper sources on eggshell quality and phosphorus retention in laying hens[J].The Journal of Applied Poultry Research,2012,21(3):460-466.

[31] ATTIA Y A,ABDALAH A A,ZEWEIL H S,et al.Effect of inorganic or organic copper additions on reproductive performance,lipid metabolism and morphology of organs of dual-purpose breeding hens[J].Archiv für Geflügelkunde,2011,75(3):169-178.

[32] SWIATKIEWICZ S,KORELESKI J.The Effect of zinc and manganese source in the diet for laying hens on eggshell and bones quality[J].Veterinarni Medicina,2008,53(10):555-563.
[33] GHEISARI A A,SANEI A,SAMIE A,et al.Effect of diets supplemented with different levels of manganese,zinc,and copper from their organic or inorganic sources on egg production and quality characteristics in laying hens[J].Biological Trace Element Research,2011,142(3):557-571.

[34] FAVERO A,VIEIRA S L,ANGEL C R,et al.Reproductive performance of Cobb 500 breeder hens fed diets supplemented with zinc,manganese,and copper from inorganic and amino acid-complexed sources[J].The Journal of Applied Poultry Research,2013,22(1):80-91.

[35] STEFANELLO C,SANTOS T C,MURAKAMI A E,et al.Productive performance,eggshell quality,and eggshell ultrastructure of laying hens fed diets supplemented with organic trace minerals[J].Poultry Science,2014,93(1):104-113.

[36] VENGLOVSKÁ K,GREŠÁKOVÁ L,PLACHÁ I,et al.Effects of feed supplementation with manganese from its different sources on performance and egg parameters of laying hens[J].Czech Journal of Animal Science,2014,59(4):147-155.
[37] MA W Q,GU Y,LU J Y,et al.Effects of chromium propionate on egg production,egg quality,plasma biochemical parameters,and egg chromium deposition in late-phase laying hens[J].Biological Trace Element Research,2014,157(2):113-119.

[38] INVERNIZZI G,AGAZZI A,FERRONI M,et al.Effects of inclusion of selenium-enriched yeast in the diet of laying hens on performance,eggshell quality,and selenium tissue deposition[J].Italian Journal of Animal Science,2013,12(1):e1.
[39] ATTIA Y A,ABDALAH A A,ZEWEIL H S,et al.Effect of inorganic or organic selenium supplementation on productive performance,egg quality and some physiological traits of dual-purpose breeding hens[J].Czech Journal of Animal Science,2010,55(11):505-519.
[40] CUFADAR Y,OLGUN O,BAHTIYARCA Y.Effects of different sources of boron supplementation to diet on egg shell quality and bone characteristics in laying hens[J].Archiva Zootechnica,2011,14(3):37-42.
[41] KIM C H,PAIK I K,KIL D Y,et al.Effects of dietary magnesium concentrations on performance and eggshell quality of laying hens[J].Journal of Animal and Veterinary Advances,2013,12(1):104-107.
[42] SWIATKIEWICZ S,ARCZEWSKA-WŁOSEK A.Prebiotic fructans and organic acids as feed additives improving mineral availability[J].World's Poultry Science Journal,2012,68(2):269-279.

[43] CHEN Y C,CHEN T C.Mineral utilization in layers as influenced by dietary oligofructose and inulin[J].International Journal of Poultry Science,2004,3(7):442-445.

[44] ŚWIATKIEWICZ S,KORELESKI J,ARCZEWSKA A.Laying performance and eggshell quality in laying hens fed diets supplemented with prebiotics and organic acids[J].Czech Journal of Animal Science,2010,55(7):294-304.
[45] ŚWIATKIEWICZ S,ARCZEWSKA-WŁOSEK A,KRAWCZYK J,et al.Effects of selected feed additives on the performance of laying hens given a diet rich in maize dried distiller's grains with solubles (DDGS)[J].British Poultry Science,2013,54(4):478-485.

[46] CESARI V,MANGIAGALLI M G,GIARDINI A,et al.Egg quality and productive performance of laying hens fed different levels of skimmed milk powder added to a diet containing
Lactobacillus acidophilus[J].Poultry Science,2014,93(5):1197-1201.

[47] YILDIZ G,SACAKLI P,GUNGOR T.The effect of dietary Jerusalem artichoke (
Helianthus tuberosus L.) on performance,egg quality characteristics and egg cholesterol content in laying hens[J].Czech Journal of Animal Science,2006,51(8):349-354.
[48] DE LA MORA L J P,OROZCO-HERNÁNDEZ J R,DE JESÙS RUÍZ-GARCÍA I,et al.Quail egg yield and quality of the
Coturnix coturnix response to the addition level of agave inulin to the drinking water[J].Italian Journal of Animal Science,2014,13(2):2981.
[49] HASHIM M,FOWLER J,HAQ A,et al.Effect of yeast cell wall on earlyproduction laying hen performance[J].The Journal of Applied Poultry Research,2013,22(4):792-797.

[50] YOUSEFI M,KARKOODI K.Effect of probiotic Thepax[XC1.TIF] and
Saccharomyces cerevisiae supplementation on performance and egg quality of laying hens[J].International Journal of Poultry Science,2007,6(1):52-54.

[51] PANDA A K,RAMA RAO S V,RAJU M V L N,et al.Effect of probiotic (
Lactobacillus sporogenes) feeding on egg production and quality,yolk cholesterol and humoral immune response of White Leghorn layer breeders[J].Journal of the Science of Food and Agriculture,2008,88(1):43-47.

[52] MIKULSKI D,JANKOWSKI J,NACZMANSKI J,et al.Effects of dietary probiotic (
Pediococcus acidilactici) supplementation on performance,nutrient digestibility,egg traits,egg yolk cholesterol,and fatty acid profile in laying hens[J].Poultry Science,2012,91(10):2691-2700.

[53] ABDELQADER A,IRSHAID R,AL-FATAFTAH A R.Effects of dietary probiotic inclusion on performance,eggshell quality,cecal microflora composition,and tibia traits of laying hens in the late phase of production[J].Tropical Animal Health and Production,2013,45(4):1017-1024.

[54] ABDELQADER A,AL-FATAFTAH A R,DA ŞG.Effects of dietary
Bacillus subtilis and inulin supplementation on performance,eggshell quality,intestinal morphology and microflora composition of laying hens in the late phase of production[J].Animal Feed Science and Technology,2013,179(1/2/3/4):103-111.
[55] SCHOLZ-AHRENS K E,SCHAAFSMA G,VAN DEN HEUVEL E G H M,et al.Effects of prebiotics on mineral metabolism[J].The American Journal of Clinical Nutrition,2001,73(2):459S-464S.
[56] YÖRVK M A,GVL M,HAYIRLI A,et al.The effects of supplementation of humate and probiotic on egg production and quality parameters during the late laying period in hens[J].Poultry Science 2004,83(1):84-88.
[57] SENGOR E,YARDIMCI M,CETINGUL S,et al.Effects of short chain fatty acid (SCFA) supplementation on performance and egg characteristics of old breeder hens[J].South African Journal of Animal Science,2007,37(3):158-163.
[58] SOLTAN M A.Effect of dietary organic acid supplementation on egg production,egg quality,and some blood serum parameters in laying hens[J].International Journal of Poultry Science,2008,7(6):613-621.

[59] YESILBAG D,ÇOLPAN I.Effects of organic acid supplemented diets on growth performance,egg production and quality and on serum parameters in laying hens[J]. Revue de Médecine Vétérinaire,2006,157(5):280-284.

[60] ÇABUK M,BOZKURT M,ALÇIÇEK A,et al.Effect of a dietary essential oil mixture on performance of laying hens in the summer season[J].South African Journal of Animal Science,2006,36(4):215-221.
[61] KAYA H,KAYA A,CELEBI S,et al.Effects of dietary supplementation of essential oils and vitamin E on performance,egg quality and
Escherichia coli count in excreta[J].Indian Journal of Animal Research,2013,47(6):515-520.
[62] LOKAEWMANEE K,YAMAUCHI K E,KOMORI T,et al.Eggshell quality,eggshell structure and small intestinal histology in laying hens fed dietary Pantoea-6[XC1.TIF] and plant extracts[J].Italian Journal of Animal Science,2014,13(2):3136.
[63] BOKA J,MAHDAVI A H,SAMIE A H,et al.Effect of different levels of black cumin (
Nigella sativa L.) on performance,intestinal
Escherichia coli colonization and jejunal morphology in laying hens[J].Journal of Animal Physiology and Animal Nutrition,2014,98(2):373-383.

[64] SHARMA R K,RAVIKANTH K,MAINI S,et al.Influence of calcium and phosphorus supplements with synergistic herbs on egg shell quality in late layers[J].Veterinary World,2009,2(6):231-233.
[65] ZHOU Z L,DENG Y F,TAO Q S,et al.Effects of Gushukang,a Chinese herbal medicine,on bone characteristics and osteoporosis in laying hens[J].Poultry Science,2009,88(11):2342-2345.

[66] BOZKURT M,KÜÇÜKYILMAZ K,PAMUKÇU M,et al.Long-term effects of dietary supplementation with an essential oil mixture on the growth and laying performance of two layer strains[J].Italian Journal of Animal Science,2012,11(1):e5.