[1] 贾玉东,雷霁霖.硬骨鱼类卵子质量研究进展[J].中国水产科学,2012,19(3):545-555.
[2] MARCH B E.Essential fatty acids in fish physiology[J].Canadian Journal of Physiology and Pharmacology,1993,71(9):684-689.

[3] IZQUIERDO M S,FERNANDEZ-PALACIOS H,TACON A G J.Effect of broodstock nutrition on reproductive performance of fish[J].Aquaculture,2001,197(1/2/3/4):25-42.
[4] ZAKERI M,KOCHANIAN P,MARAMMAZI J G,et al.Effects of dietary n-3 HUFA concentrations on spawning performance and fatty acids composition of broodstock,eggs and larvae in yellowfin sea bream,
Acanthopagrus latus[J].Aquaculture,2011,310(3/4):388-394.
[5] TOCHER D R.Fatty acid requirements in ontogeny of marine and freshwater fish[J].Aquaculture Research,2010,41(5):717-732.

[6] 许友卿,刘晓丽,安晓玲,等.高度不饱和脂肪酸对水生动物代谢的影响及其机理[J].饲料工业,2017,38(4):1-7.
[7] 吉红,田晶晶.高不饱和脂肪酸(HUFAs)在淡水鱼类中的营养作用研究进展[J].水产学报,2014,38(9):1650-1665.
[8] 谭青松,吴凡,杜浩,等.饲料营养对亲鱼生殖性能的影响研究进展[J].水生态学杂志,2016,37(4):1-9.
[9] DA SILVA F F G,STØTTRUP J G,KJØRSVIK E,et al.Interactive effects of dietary composition and hormonal treatment on reproductive development of cultured female European eel,
Anguilla Anguilla[J].Animal Reproduction Science,2016,171:17-26.
[10] BAEZA R,PEÑARANDA D S,VÍLCHEZ M C,et al.Exploring correlations between sex steroids and fatty acids and their potential roles in the induced maturation of the male European eel[J].Aquaculture,2015,435:328-335.
[11] TURCHINI G M,FRANCIS D S,KEAST R S J,et al.Transforming salmonid aquaculture from a consumer to a producer of long chain omega-3 fatty acids[J].Food Chemistry,2011,124(2):609-614.

[12] TOCHER D R.Omega-3 long-chain polyunsaturated fatty acids and aquaculture in perspective[J].Aquaculture,2015,449:94-107.
[13] VAGNER M,SANTIGOSA E.Characterization and modulation of gene expression and enzymatic activity of delta-6 desaturase in teleosts:a review[J].Aquaculture,2011,315(1/2):131-143.
[14] EMERY J A,HERMON K,HAMID N K,et al.△-6 desaturase substrate competition:dietary linoleic acid (18:2n-6) has only trivial effects on α-linolenic acid (18:3n-3) bioconversion in the teleost rainbow trout[J].PLoS One,2013,8(2):e57463.
[15] SPRECHER H.Metabolism of highly unsaturated n-3 and n-6 fatty acids[J].Biochimica et Biophysica:Molecular and Cell Biology of Lipids,2000,1486(2/3):219-231.
[16] LI Y Y,MONROIG O,ZHANG L,et al.Vertebrate fatty acyl desaturase with △4 activity[J].Proceedings of the National Academy of Sciences of the United States of America,2010,107(39):16840-16845.

[17] MORAIS S,CASTANHEIRA F,MARTINEZ-RUBIO L,et al.Long chain polyunsaturated fatty acid synthesis in a marine vertebrate:ontogenetic and nutritional regulation of a fatty acyl desaturase with △4 activity[J].Biochimica et Biophysica Acta:Molecular and Cell Biology of Lipids,2012,1821(4):660-671.

[18] FONSECA-MADRIGAL J,NAVARRO J C,HONTORIA F,et al.Diversification of substrate specificities in teleostei Fads2:characterization of △4 and △6△5 desaturases of
Chirostoma estor[J].Journal of Lipid Research,2014,55(7):1408-1419.

[19] KUAH M K,JAYA-RAM A,SHU-CHIEN A C.The capacity for long-chain polyunsaturated fatty acid synthesis in a carnivorous vertebrate:functional characterisation and nutritional regulation of a Fads2 fatty acyl desaturase with △4 activity and an Elovl5 elongase in striped snakehead (
Channa striata)[J].Biochimica et Biophysica:Molecular and Cell Biology of Lipids,2015,1851(3):248-260.
[20] 谢帝芝,陈芳,张庆昊,等.鱼类LC-PUFA合成代谢调控机制研究进展[J].汕头大学学报(自然科学版),2015,30(2):3-19.
[21] JAYA-RAM A,KUAH M K,LIM P S,et al.Influence of dietary HUFA levels on reproductive performance,tissue fatty acid profile and desaturase and elongase mRNAs expression in female zebrafish
Danio rerio[J].Aquaculture,2008,277(3/4):275-281.
[22] 郭盼,周继术,吉红,等.不同链长脂肪酸对斑马鱼的性腺脂肪酸组成、繁殖力与仔鱼成活率的影响[J].水生生物学报,2017,41(4):766-773.
[23] LUO L,AI L C,LI T L,et al.The impact of dietary DHA/EPA ratio on spawning performance,egg and offspring uality in Siberian sturgeon (
Acipenser baeri)[J].Aquaculture,2015,437:140-145.
[24] 齐茜,刘晓勇,孙大江,等.饲料脂肪源搭配对施氏鲟雌鱼繁殖性能和血清性类固醇激素的影响[J].中国水产科学,2014,21(1):118-124.
[25] KÖPRVCV K,YONAR M E,ÖZCAN S.Effect of dietary n-3 polyunsaturated fatty acids on antioxidant defense and sperm quality in rainbow trout (
Oncorhynchus mykiss) under regular stripping conditions[J].Animal Reproduction Science,2015,163:135-143.
[26] EL-SAYED A F M,MANSOUR C R,EZZAT A A.Effects of dietary lipid source on spawning performance of Nile tilapia (
Oreochromis niloticus) broodstock reared at different water salinities[J].Aquaculture,2005,248(1/2/3/4):187-196.
[27] CALLAN C K,LAIDLEY C W,KLING L J,et al.The effects of dietary HUFA level on flame angelfish (
Centropyge loriculus) spawning,egg quality and early larval characteristics[J].Aquaculture Research,2014,45(7):1176-1186.

[28] FURUITA H,TANAKA H,YAMAMOTO T et al.Effects of high levels of n-3 HUFA in broodstock diet on egg quality and egg fatty acid composition of Japanese flounder,
Paralichthys olivaceus[J].2002,Aquaculture,210(1/2/3/4):323-333.
[29] FERNÁNDEZ-PALACIOS H,IZQUIERDO M S,ROBAINA L,et al.Effect of n-3 HUFA level in broodstock diets on egg quality of gilthead sea bream (
Sparus aurata L.)[J].Aquaculture,1995,132(3/4):325-337.
[30] LI Y Y,CHEN W Z,SUN Z W,et al.Effects of n-3 HUFA content in broodstock diet on spawning performance and fatty acid composition of eggs and larvae in
Plectorhynchus cinctus[J].Aquaculture,2005,245(1/2/3/4):263-272.
[31] 温海深,林浩然.环境因子对硬骨鱼类性腺发育成熟及其排卵和产卵的调控[J].应用生态学报,2001,12(1):151-155.
[32] LUBZENS E,YOUNG G,BOBE J,et al.Oogenesisin teleosts:how fish eggs are formed[J].General and Comparative Endocrinology,2010,165(3):367-389.

[33] BRANSDEN M P,BATTAGLENE S C,GOLDSMID R M,et al.Broodstock condition,egg morphology and lipid content and composition during the spawning season of captive striped trumpeter,
Latris lineate[J].Aquaculture,2007,268(1/2/3/4):2-12.
[34] 彭士明,李云莉,施兆鸿,等.海水鱼类亲体必需脂肪酸营养的研究进展[J].海洋渔业,2016,38(1):98-106.
[35] 李远友,陈伟洲,孙泽伟,等.饲料中n-3 HUFA含量对花尾胡椒鲷亲鱼的生殖性能及血浆性类固醇激素水平季节变化的影响[J].动物学研究,2004,25(3):249-255.
[36] PENG S M,GAO Q X,SHI Z H,et al.Effect of dietary n-3 LC-PUFAs on plasma vitellogenin,sex steroids,and ovarian steroidogenesis during vitellogenesis in female silver pomfret (
Pampus argenteus) broodstock[J].Aquaculture,2015,444:93-98.
[37] DUPONT J,REVERCHON M,BERTOLO M J,et al.Nutritional signals and reproduction[J].Molecular and Cellular Endocrinology,2014,382(1):527-537.

[38] IFFO.The marine ingredients organisation:Global standard for responsible supply (IFFO-RS)[J/OL].2012.http://www.iffo.net/.
[39] AFONSO C,CARDODO C,FREIRE M,et al.The impact of alternative dietary lipids on the
in vitro bioaccessibility of sole fillets for human consumption[J].Aquaculture,2017,474(1):66-74.
[40] IZQUIERDO M S,TURKMEN S,MONTERO D,et al.Nutritional programming through broodstock diets to improve utilization of very low fishmeal and fish oil diets in gilthead sea bream[J].Aquaculture,2015,449:18-26.
[41] 潘瑜,陈效儒,马霞,等.裂殖壶菌替代鱼油对海鲈生长和抗氧化能力的影响[J].饲料工业,2016,37(4):24-27.
[42] LI M H,ROBINSON E H,TUCKER C S,et al.Effects of dried algae
Schizochytrium sp.,a rich source of docosahexaenoic acid,on growth,fatty acid composition,and sensory quality of channel catfish
Ictalurus punctatus[J].Aquaculture,2009,292(3/4):232-236.
[43] 周俊宇,杨志刚.富含HUFA海洋微藻在水产饲料中的应用[J].饲料研究,2017(7):40-44.