[1] SAPER C B,CHOU T C,ELMQUISTl J K.The need to feed:homeostatic and hedonic control of eating[J].Neuron,2002,36(2):199-211.

[2] 田娟,何艮,麦康森,等.鱼类食欲调控研究进展[J].动物营养学报,2016,28(4):984-998.
[3] VOLKOFF H.The neuroendocrine regulation of food intake in fish:a review of current knowledge[J].Frontiers in Neuroscience,2016,10:540.
[4] VALE W,SPIESS J,RIVIER C,et al.Characterization of a 41-residue ovine hypothalamic peptide that stimulates secretion of corticotropin and beta-endorphin[J].Science,1981,213(4514):1394-1397.

[5] SAFFRAN M,SCHALLY A V.The release of corticotrophin by anterior pituitary tissue
in vitro[J].Canadian Journal of Biochemistry and Physiology,1955,33(3):408-415.

[6] FURUTANI Y,MORIMOTO Y,SHIBAHARA S,et al.Cloning and sequence analysis of cDNA for ovine corticotropin-releasing factor precursor[J].Nature,1983,301(5900):537-540.

[7] VANDENBORNE K,DE GROEF B,GEELISSEN S M,et al.Molecular cloning and developmental expression of corticotropin-releasing factor in the chicken[J].Endocrinology,2005,146(1):301-308.

[8] OKAWARA Y,MORLEY S D,BURZIO L O,et al.Cloning and sequence analysis of cDNA for corticotropin-releasing factor precursor from the teleost fish
Catostomus commersoni[J].Proceedings of the National Academy of Sciences of the United States of America,1988,85(22):8439-8443.

[9] BERNIER N J,LIN X W,PETER R E.Differential expression of corticotropin-releasing factor (CRF) and urotensin Ⅰ precursor genes,and evidence of
CRF gene expression regulated by cortisol in goldfish brain[J].General and Comparative Endocrinology,1999,116(3):461-477.

[10] ANDO H,HASEGAWA M,ANDO J,et al.Expression of salmon corticotropin-releasing hormone precursor gene in the preoptic nucleus in stressed rainbow trout[J].General and Comparative Endocrinology,1999,113(1):87-95.

[11] DOYON C,GILMOUR K,TRUDEAU V,et al.Corticotropin-releasing factor and neuropeptide Y mRNA levels are elevated in the preoptic area of socially subordinate rainbow trout[J].General and Comparative Endocrinology,2003,133(2):260-271.

[12] HUISING M O,METZ J R,VAN SCHOOTEN C,et al.Structural characterisation of a cyprinid (
Cyprinus carpio L.) CRH,CRH-BP and CRH-R1,and the role of these proteins in the acute stress response[J].Journal of Molecular Endocrinology,2004,32(3):627-648.

[13] VAN ENCKEVORT F H J,PEPELS P P L M,LEUNISSEN J A M,et al.
Oreochromis mossambicus (tilapia) corticotropin-releasing hormone:cDNA sequence and bioactivity[J].Journal of Neuroendocrinology,2000,12(2):177-186.
[14] LU W Q,DOW L,GUMUSGOZ S,et al.Coexpression of corticotropin-releasing hormone and urotensin Ⅰ precursor genes in the caudal neurosecretory system of the euryhaline flounder (
Platichthys flesus):a possible shared role in peripheral regulation[J].Endocrinology,2004,145(12):5786-5797.

[15] CHANDRASEKAR G,LAUTER G,HAUPTMANN G.Distribution of corticotropin-releasing hormone in the developing zebrafish brain[J].Journal of Comparative Neurology,2007,505(4):337-351.

[16] WANG T,ZHOU C W,YUAN D Y,et al.
Schizothorax prenanti corticotropin-releasing hormone (CRH):molecular cloning,tissue expression,and the function of feeding regulation[J].Fish Physiology and Biochemistry,2014,40(5):1407-1415.

[17] MATSUDA K,MORIMOTO N,HASHIMOTO K,et al.Changes in the distribution of corticotropin-releasing factor (CRF)-like immunoreactivity in the larval bullfrog brain and the involvement of CRF in the cessation of food intake during metamorphosis[J].General and Comparative Endocrinology,2010,168(2):280-286.

[18] MATSUDA K.Regulation of feeding behavior and psychomotor activity by corticotropin-releasing hormone (CRH) in fish[J].Frontiers in Neuroscience,2013,7:91.
[19] DE PEDRO N,ALONSO-GÓMEZ A L,GANCEDO B,et al.Role of corticotropin-releasing factor (CRF) as a food intake regulator in goldfish[J].Physiology & Behavior,1993,53(3):517-520.

[20] BERNIER N J,PETER R E.Appetite-suppressing effects of urotensin Ⅰ and corticotropin-releasing hormone in goldfish (Carassius auratus)[J].Neuroendocrinology,2001,73(4):248-260.

[21] PARROTT R.Central administration of corticotropin releasing factor in the pig:effects on operant feeding,drinking and plasma cortisol[J].Physiology & Behavior,1990,47(3):519-524.

[22] YUAN P Q,WU S V,ELLIOTT J,et al.Expression of corticotropin releasing factor receptor type 1(CRF 1) in the human gastrointestinal tract and upregulation in the colonic mucosa in patients with ulcerative colitis[J].Peptides,2012,38(1):62-69.

[23] COLOMBO E,SANGIOVANNI E,DELL'AGLI M.A review on the anti-inflammatory activity of pomegranate in the gastrointestinal tract[J].Evidence-Based Complementary and Alternative Medicine,2013,2013:247145.
[24] TACHÉ Y,PERDUE M H.Role of peripheral CRF signalling pathways in stress-related alterations of gut motility and mucosal function[J].Neurogastroenterology & Motility,2004,16(Suppl.1):137-142.
[25] PEPELS P P,VAN Helvoort H,Bonga S W E,et al.Corticotropin-releasing hormone in the teleost stress response:rapid appearance of the peptide in plasma of tilapia (
Oreochromis mossambicus)[J].Journal of Endocrinology,2004,180(3):425-438.

[26] YADAWA A K,CHATURVEDI C M.Expression of stress hormones AVP and CRH in the hypothalamus of
Mus musculus following water and food deprivation[J].General and Comparative Endocrinology,2016,239:13-20.
[27] MARUYAMA K,MIURA T,UCHIYAMA M,et al.Relationship between anorexigenic action of pituitary adenylate cyclase-activating polypeptide (PACAP) and that of corticotropin-releasing hormone (CRH) in the goldfish,
Carassius auratus[J].Peptides,2006,27(7):1820-1826.

[28] BERNIER N J,CRAIG P M.CRF-related peptides contribute to stress response and regulation of appetite in hypoxic rainbow trout[J].American Journal of Physiology-Regulatory,Integrative and Comparative Physiology,2005,289(4):R982-R990.
[29] WOOD C M.Dogmas and controversies in the handling of nitrogenous wastes:is exogenous ammonia a growth stimulant in fish?[J].Journal of Experimental Biology,2004,207(12):2043-2054.

[30] ORTEGA V A,RENNER K J,BERNIER N J.Appetite-suppressing effects of ammonia exposure in rainbow trout associated with regional and temporal activation of brain monoaminergic and CRF systems[J].Journal of Experimental Biology,2005,208(10):1855-1866.

[31] CRAIG P M,AL-TIMIMI H,ERNIER N J.Differential increase in forebrain and caudal neurosecretory system corticotropin-releasing factor and urotensin Ⅰ gene expression associated with seawater transfer in rainbow trout[J].Endocrinology,2005,146(9):3851-3860.

[32] VAUGHAN J,DONALDSON C,BITTENCOURT J,et al.Urocortin,a mammalian neuropeptide related to fish urotensin Ⅰ and to corticotropin-releasing factor[J].Nature,1995,378(6554):287-292.

[33] BERNIER N J,PETER R E.The hypothalamic-pituitary-interrenal axis and the control of food intake in teleost fish[J].Comparative Biochemistry and Physiology Part B:Biochemistry and Molecular Biology,2001,129(2/3):639-644.
[34] BERTHOUD H R.Multiple neural systems controlling food intake and body weight[J].Neuroscience & Biobehavioral Reviews,2002,26(4):393-428.

[35] ARAI M,ASSIL I Q,ABOU-SAMRA A B.Characterization of three corticotropin-releasing factor receptors in catfish:a novel third receptor is predominantly expressed in pituitary and urophysis[J].Endocrinology,2001,142(1):446-454.

[36] CARDOSO J C R,POWER D M,ELGA R,et al.Isolation and characterisation of the corticotropin releasing factor receptor 1(
CRFR1) gene in a teleost fish,
Fugu rubripes[J].DNA Sequence,2003,14(3):215-218.

[37] MARTÍNEZ V,WANG L X,RIVIER J E,et al.Differential actions of peripheral corticotropin-releasing factor (CRF),urocortinⅡ,and urocortin Ⅲ on gastric emptying and colonic transit in mice:role of CRF receptor subtypes 1 and 2[J].Journal of Pharmacology and Experimental Therapeutics,2002,301(2):611-617.

[38] GREGORY T R,WOOD C M.The effects of chronic plasma cortisol elevation on the feeding behaviour,growth,competitive ability,and swimming performance of juvenile rainbow trout[J].Physiological and Biochemical Zoology,1999,72(3):286-295.

[39] BERNIER N J,BEDARD N,PETER R E.Effects of cortisol on food intake,growth,and forebrain neuropeptide Y and corticotropin-releasing factor gene expression in goldfish[J].General and Comparative Endocrinology,2004,135(2):230-240.

[40] DUCOURET B,TUJAGUE M,ASHRAF J,et al.Cloning of a teleost fish glucocorticoid receptor shows that it contains a deoxyribonucleic acid-binding domain different from that of mammals[J].Endocrinology,1995,136(9):3774-3783.

[41] VLLETTE P A,YOUNG G.Tissue culture of sockeye salmon intestine:functional response of Na
+-K
+-ATPase to cortisol[J].American Journal of Physiology-Regulatory,Integrative and Comparative Physiology,2005,288(6):R1598-R1605.
[42] SMITH S M,VAUGHAN J M,DONALDSON C J,et al.Cocaine-and amphetamine-regulated transcript activates the hypothalamic-pituitary-adrenal axis through a corticotropin-releasing factor receptor-dependent mechanism[J].Endocrinology,2004,145(11):5202-5209.

[43] HEINRICHS S C,MENZAGHI F,PICH E M,et al.Corticotropin-releasing factor in the paraventricular nucleus modulates feeding induced by neuropeptide Y[J].Brain Research,1993,611(1):18-24.

[44] BAZHAN N,ZELENA D.Food-intake regulation during stress by the hypothalamo-pituitary-adrenal axis[J].Brain Research Bulletin,2013,95:46-53.
[45] MATSUDA K,KOJIMA K,SHIMAKURA S I,et al.Corticotropin-releasing hormone mediates α-melanocyte-stimulating hormone-induced anorexigenic action in goldfish[J].Peptides,2008,29(11):1930-1936.

[46] SOLOMON A,DE FANTI B A,MARTÍNEZ A M.Peripheral ghrelin participates in glucostatic feeding mechanisms and in the anorexigenic signalling mediated by CART and CRF neurons[J].Nutritional Neuroscience,2005,8(5/6):287-295.
[47] JÖNSSON E,KAIYA H,BJÖRNSSON B T.Ghrelin decreases food intake in juvenile rainbow trout (
Oncorhynchus mykiss) through the central anorexigenic corticotropin-releasing factor system[J].General and Comparative Endocrinology,2010,166(1):39-46.

[48] TAHERI S,MURPHY K,COHEN M,et al.The effects of centrally administered apelin-13 on food intake,water intake and pituitary hormone release in rats[J].Biochemical and Biophysical Research Communications,2002,291(5):1208-1212.

[49] LV S Y,YANG Y J,QIN Y J,et al.Central apelin-13 inhibits food intake via the CRF receptor in mice[J].Peptides,2012,33(1):132-138.