[1] 郭宝林,贾志海,张玉枝.畜禽铜营养研究进展[J].饲料工业,2005,26(10):48-52. GUO B L,JIA Z H,ZHANG Y Z.Research progress of copper nutrition in livestock and poultry[J].Feed Industry,2005,26(10):48-52.(in Chinese)
[2] 刘培培.铜在家禽营养中的应用[J].饲料博览,2015(9):41-43. LIU P P.The application of copper in poultry nutrition[J].Feed Review,2015(9):41-43.(in Chinese)
[3] 王丽赛,张丽阳,邵玉新,等.我国畜禽饲料资源中微量元素铜含量分布的调查[J].中国农业科学,2019,52(11):1982-1992. WANG L S,ZHANG L Y,SHAO Y X,et al.A survey on distribution of copper contents in feedstuffs for livestock and poultry in China[J].Scientia Agricultura Sinica,2019,52(11):1982-1992.(in Chinese)
[4] HARRIS E D.Progress in clinical & bioloical research[J].The Transport of Copper,1993,380:163-179.
[5] HELLMAN N E,GITLIN J D.Ceruloplasmin metabolism and function[J].Annual Review of Nutrition,2002,22:439-458.
[6] GOODMAN V L,BREWER G J,MERAJVER S D.Copper deficiency as an anti-cancer strategy[J].Endocrine-Related Cancer,2004,11(2):255-263.

[7] 吴学壮,闻治国,胡洪,等.斜率比法评定肉仔鸡对铜源的相对生物学利用率[J].动物营养学报,2019,31(4):1596-1603. WU X Z,WEN Z G,HU H,et al.Evaluation of relative bioavailability of copper source with slope ratio method for broilers[J].Chinese Journal of Animal Nutrition,2019,31(4):1596-1603.(in Chinese)
[8] KAYA A,ALTINER A,ÖZPINAR A.Effect of copper deficiency on blood lipid profile and haematological parameters in broilers[J].Transboundary and Emerging Diseases,2010,53(8):399-404.
[9] WU X Z,DAI S F,HUA J L,et al.Influence of dietary copper methionine concentrations on growth performance,digestibility of nutrients,serum lipid profiles,and immune defenses in broilers[J].Biological Trace Element Research,2019,191(1):199-206.

[10] OGNIK K,TUTAJ K,CHOLEWIŃSKA E,et al.The effect of a rat diet without added Cu on redox status in tissues and epigenetic changes in the brain[J].Annals of Animal Science,2020,20(2):503-520.

[11] GAETKE L M,CHOW-JOHNSON H S,CHOW C K.Copper:toxicological relevance and mechanisms[J].Archives of Toxicology,2014,88(11):1929-1938.

[12] 魏晓玫,李浩棠,郭小权,等.日粮硫酸铜对育成蛋鸡血清铜、肝铜含量及相关肝酶的影响[J].养禽与禽病防治,2009(1):14-17. WEI X M,LI H T,GUO X Q,et al.Effects of dietary copper sulfate on serum copper,liver copper content and related liver enzymes in laying hens[J].Poultry Husbandry and Disease Control,2009(1):14-17.(in Chinese)
[13] BOZKAYA L A,OZTÜRK-UREK R,AYDEMIR T,et al.Effects of Se,Cu and Se+vitamin E deficiency on the activities of CuZnSOD,GSH-Px,CAT and LPO levels in chicken erythrocytes[J].Cell Biochemistry and Function,2010,19(3):153-157.
[14] 胡锴,任常宝,颜城,等.高铜日粮对肉鸡肝脏抗氧化功能的影响[J].中国家禽,2011,33(12):26-29. HU K,REN C B,YAN C,et al.Effect of different levels of dietary copper sulfate on liver antioxidantal index in broilers[J].China Poultry,2011,33(12):26-29.(in Chinese)
[15] HAWK S N,LANOUE L,KEEN C L,et al.Copper-deficient rat embryos are characterized by low superoxide dismutase activity and elevated superoxide anions[J].Biology of Reproduction,2003,68(3):896-903.

[16] MAJEWSKI M,OGNIK K,ZDUNCZYK P,et al.Effect of dietary copper nanoparticles versus one copper (Ⅱ) salt:analysis of vasoreactivity in a rat model[J].Pharmacological Reports,2017,69(6):1282-1288.

[17] ANGELOVA M,ASENOVA S,NEDKOVA V,et al.Copper in the human organism[J].Trakia Journal of Sciences,2011,9(1):88-98.
[18] GUO R,HENRY P R,HOLWERDA R A,et al.Chemical characteristics and relative bioavailability of supplemental organic copper sources for poultry[J].Journal of Animal Science,2001,79(5):1132-1141.

[19] KIM J W,KIL D Y.Determination of relative bioavailability of copper in tribasic copper chloride to copper in copper sulfate for broiler chickens based on liver and feather copper concentrations[J].Animal Feed Science and Technology,2015,210:138-143.
[20] LEDOUX D R,HENRY P R,AMMERMAN C B,et al.Estimation of the relative bioavailability of inorganic copper sources for chicks using tissue uptake of copper[J].Journal of Animal Science,1991,69(1):215-222.

[21] LUO X G,JI F,LIN Y X,et al.Effects of dietary supplementation with copper sulfate or tribasic copper chloride on broiler performance,relative copper bioavailability,and oxidation stability of vitamin E in feed[J].Poultry Science,2005,84(6):888-893.

[22] 杨凤.动物营养学[M].北京:农业出版社,1993:67-68. YANG F.Animal nutrition[M].Beijing:China Agricultural Press,1993,67-68.(in Chinese)
[23] 呙于明.家禽营养与饲料[M].北京:中国农业大学出版社,1997:123-124. GUO Y M.Poultry nutrition and feed[M].Beijing:China Agricultural University Press,1997:123-124.(in Chinese)
[24] 马雪莲,张丽阳,王良治,等.我国畜禽饲料资源中微量元素铁含量分布的调查研究[J].动物营养学报,2019,31(8):3821-3829. MA X L,ZHANG L Y,WANG L Z,et al.A survey on distribution of iron contents in feedstuffs for livestock and poultry in China[J].Chinese Journal of Animal Nutrition,2019,31(8):3821-3829.(in Chinese)
[25] 林丽,林萍.地中海贫血和缺铁性贫血诊断中血常规检验的应用效果观察[J].中外医学研究,2019,17(2):62-64. LIN L,LIN P.Observation on the application effect of blood routine test in the diagnosis of thalassemia and iron deficiency anemia[J].Chinese and Foreign Medical Research,2019,17(2):62-64.(in Chinese)
[26] MA X Y,LIU S B,LU L,et al.Relative bioavailability of iron proteinate for broilers fed a casein-dextrose diet[J].Poultry Science,2014,93(3):556-563.

[27] 孙健.甘氨酸亚铁对肉仔鸡生理生化指标及相关基因表达量的影响[D].硕士学位论文.哈尔滨:东北农业大学,2015. SUN J.Effects of iron glycine on physiological and biochemical indicators and related gene expression in broiler chickens[D].Master's Thesis.Harbin:Northeast Agricultural University,2015. (in Chinese)
[28] 张伶燕.有机铁源的化学特性、对肉仔鸡的相对生物学利用率及其小肠铁吸收研究[D].博士学位论文.北京:中国农业科学院,2016. ZHANG L Y.Study on chemical characteristics,relative bioavailabilites and small intestinal iron absorptions of organic iron sources in broilers[D].Ph.D.Thesis.Beijing:Chinese Academy of Agricultural Sciences,2016.(in Chinese)
[29] 马新燕.肉仔鸡对有机蛋白铁相对生物学利用率及其饲粮铁适宜水平的研究[D]. 硕士学位论文.北京:中国农业科学院,2012. MA X Y.Study on relative bioavailabilitiy of organic protein iron source and optimal dietary iron level for broilers[D]. Master's Thesis.Beijing:Chinese Academy of Agricultural Sciences,2012.(in Chinese)
[30] KOHEN R,NYSKA A.Oxidation of biological systems:oxidative stress phenomena,antioxidants,redox reactions,and methods for their quantification[J].Toxicologic Pathology,2002,30(6):620-650.

[31] 吴建设,呙于明,周毓平.日粮铁缺乏对肉仔鸡机体抗氧化功能影响的研究[J].中国畜牧杂志,1999,35(4):5-7. WU J S,GUO Y M,ZHOU Y P.Effects of dietary iron on anti-oxidative function in broiler chick[J].Chinese Journal of Animal Science,1999,35(4):5-7.(in Chinese)
[32] 马春艳.22-42日龄肉鸡玉米-豆粕型饲粮铁适宜水平的研究[D].硕士学位论文.北京:中国农业科学院,2014. MA C Y.Study on an optimal dietary iron level for broilers fed a corn-soybean meal diet from 22 to 42 days of age[D].Master's Thesis.Beijing:Chinese Academy of Agricultural Sciences,2014.(in Chinese)
[33] 贾彦芳.肉仔鸡对几种铁源吸收的研究[D].硕士学位论文.哈尔滨:东北农业大学,2015. JIA Y F.Studies on absorption of iron sources in the broiler[D].Master's Thesis.Harbin:Northeast Agricultural University,2015.(in Chinese)
[34] 侯韬,徐燕平.缺铁性贫血患者检测血清铁和转铁蛋白的临床意义[J].中国现代医生,2010,48(29):155-156. HOU T,XU Y P.Clinical significance of detection of serum iron and transferrin in patients with iron deficiency anemia[J].China Modern Doctor,2010,48(29):155-156.(in Chinese)
[35] 唐胜球,董小英,邹晓庭,等.不同铁源对鸡蛋铁含量的影响[J].江苏农业学报,2004,20(4):264-267. TANG S Q,DONG X Y,ZOU X T,et al.Effects of iron of different sources on iron content of egg[J].Jiangsu Journal of Agricultural Sciences,2004,20(4):264-267.(in Chinese)
[36] BEGUIN Y.Soluble transferrin receptor for the evaluation of erythropoiesis and iron status[J].Clinica Chimica Acta,2003,329(1/2):9-22.
[37] 周桂莲,韩友文,徐子伟,等.不同铁源生物学效价的研究[J].动物营养学报,2002,14(2):47-52,56. ZHOU G L,HAN Y W,XU Z W,et al.Bioavailability of iron in different fe sources for rats[J].Chinese Journal of Animal Nutrition,2002,14(2):47-52,56.(in Chinese)
[38] SHINDE P L,INGALE S L,CHOI J Y,et al.Efficiency of inorganic and organic iron sources under iron depleted conditions in broilers[J].British Poultry Science,2011,52(5):578-583.

[39] CAO J,LUO X G,HENRY P R,et al.Effect of dietary iron concentration,age,and length of iron feeding on feed intake and tissue iron concentration of broiler chicks for use as a bioassay of supplemental iron sources[J].Poultry Science,1996,75(4):495-504.

[40] 郭荣富,陈克嶙,张曦,等.肉鸡蛋氨酸铁生物利用率试验研究[J].中国家禽,2004,8(增刊1):93-95. GUO R F,CHEN K L,ZHANG X,et al.Study on relative bioavailability of iron methionine for the broiler[J].China Poultry,2004,8(Suppl.1):93-95.(in Chinese)
[41] SEO S H,LEE H K,LEE W S,et al.The effect of level and period of Fe-methionine chelate supplementation on the iron content of boiler meat[J].Asian-Australasian Journal of Animal Sciences,2008,21(10):1501-1505.

[42] 姜俊芳,许梓荣,张春善,等.肉仔鸡铁营养状况的标识[J].中国兽医学报,2005,25(1):101-103. JIANG J F,XU Z R,ZHANG C S,et al.Identifiers of the iron nutrition status of broilers[J].Chinese Journal of Veterinary,2005,25(1):101-103.(in Chinese)
[43] 王伟,崔连成.蛋鸡锰缺乏症的诊断及防治[J].畜牧兽医科技信息,2017(3):120. WANG W,CUI L C.Diagnosis and prevention of manganese deficiency in laying hens[J].Chinese Journal of Animal Husbandry and Veterinary Medicine,2017(3):120.(in Chinese)
[44] 王照军.锰缺乏对肉仔鸡骨骼发育相关血清学指标的影响[D].硕士学位论文.泰安:山东农业大学,2012. WANG Z J.Effects of manganese deficiency on serum biochemical index of bone metabolism in chicken[D].Master's Thesis.Taian:Shandong Agricultural University,2012.(in Chinese)
[45] QIN S Z,LIAO X D,LU L,et al.Manganese enhances the expression of the manganese superoxide dismutase in cultured primary chick embryonic myocardial cells[J].Journal of Integrative Agriculture,2017,16(9):2038-2046.

[46] 罗绪刚,苏琪,黄俊纯,等.实用饲粮锰缺乏对肉仔鸡组织含锰超氧化歧化酶活性及其线粒体超微结构的影响[J].畜牧兽医学报,1992,23(2):97-101. LUO X G,SU Q,HUANG J C,et al.Effects of manganese (Mn) deficiency on tissue Mn containing superoxide dismutase (MnSOD) activity and its mitochondrial ultrastructures of broiler chicks fed a practical diet[J].Chinese Journal of Animal and Veterinary Sciences,1992,23(2):97-101.(in Chinese)
[47] GAO T Q,WANG F N,LI S F,et al.Manganese regulates manganese-containing superoxide dismutase (
MnSOD) expression in the primary broiler myocardial cells[J].Biological Trace Element Research,2011,144(1/2/3):695-704.
[48] LI S F,LU L,HAO S F,et al.Dietary manganese modulates expression of the manganese-containing superoxide dismutase gene in chickens[J].The Journal of Nutrition,2011,141(2):189-194.

[49] ZHAO F L,HE C,PENG H W,et al.Relative bioavailability of humate-manganese complex for broilers fed a corn-soya bean meal diet[J].Journal of Animal Physiology and Animal Nutrition,2019,103(1):108-115.

[50] XIAO J F,ZHANG Y N,WU S G,et al.Manganese supplementation enhances the synthesis of glycosaminoglycan in eggshell membrane:a strategy to improve eggshell quality in laying hens[J].Poultry Science,2014,93(2):380-388.

[51] ZHU Y W,XIE J J,LI W X,et al.Effects of environmental temperature and dietary manganese on egg production performance,egg quality,and some plasma biochemical traits of broiler breeders[J].Journal of Animal Science,2015,93(7):3431-3440.

[52] ZHANG Y N,WANG J,ZHANG H J,et al.Effect of dietary supplementation of organic or inorganic manganese on eggshell quality,ultrastructure,and components in laying hens[J].Poultry Science,2017,96(7):2184-2193.

[53] 刘彩霞.饲料锰生物有效性的评定[J].四川畜牧兽医,1999,26(12):27-28. LIU C X.Assessing bioavailability of manganese in the feedstaff[J].Sichuan Animal and Veterin Sciences,1999,26(12):27-28.(in Chinese)
[54] 罗绪刚,苏琪,黄俊纯,等.饲粮中锰(Mn)的缺乏与过量对肉仔鸡性能的长期影响[J].中国畜牧杂志,1992,28(1):11-14. LUO X G,SU Q,HUANG J C,et al.The long-term effects of manganese (Mn) deficiency and excess in a practical diet on performances of broiler chicks[J].Chinese Journal of Animal Science,1992,28(1):11-14. (in Chinese)
[55] JASEK A,COUFAL C D,PARR T M,et al.Evaluation of increasing manganese hydroxychloride level on male broiler growth performance and tibia strength[J].Journal of Applied Poultry Research,2019,28(4):1039-1047.

[56] MWANGI S,TIMMONS J,AO T,et al.Effect of manganese preconditioning and replacing inorganic manganese with organic manganese on performance of male broiler chicks[J].Poultry Science,2019,98(5):2105-2113.

[57] 罗绪刚,苏琪,黄俊纯,等.实用饲粮锰(Mn)的缺乏与过量对肉仔鸡组织中其它矿物质微量元素含量的影响[J].畜牧与兽医,1991,23(4):146-147. LUO X G,SU Q,HUANG J C,et al.The effect of deficiency and excess of manganese (Mn) in practical diets on the content of other mineral trace elements in broiler tissues[J].Animal Husbandry & Veterinary Medicine,1991,23(4):146-147.(in Chinese)
[58] PRASAD A S,MIALE A,Jr,FARID Z,et al.Zinc metabolism in patients with the syndrome of iron deficiency anemia,hepatosplenomegaly,dwarfism,and hypognadism[J].Journal of Laboratory and Clinical Medicine,1963,61:537-549.
[59] ZHANG Y N,ZHANG H J,WANG J,et al.Effect of dietary supplementation of organic or inorganic zinc on carbonic anhydrase activity in eggshell formation and quality of aged laying hens[J].Poultry Science,2017,96(7):2176-2183.

[60] 张亚男.饲粮锌对产蛋后期蛋鸡蛋壳品质及抗氧化机能的影响[D].硕士学位论文.北京:中国农业科学院,2013. ZHANG Y N.Effects of dietary zinc on eggshell quality and antioxidant status of old laying hens[J].Master's Thesis.Beijing:Chinese Academy of Agricultural Sciences,2013.(in Chinese)
[61] 孙秋娟,呙于明,张天国,等.羟基蛋氨酸螯合铜/锰/锌对产蛋鸡蛋壳品质、酶活及微量元素沉积的影响[J].中国农业大学学报,2011,16(4):127-133. SUN Q J,GUO Y M,ZHANG T G,et al.Effects of 2-hydroxy-4-(methylthio)-butanoic acid chelated Cu/Mn/Zn on eggshell quality,enzyme activity and trace mineral retention in laying hens[J].Journal of China Agricultural University,2011,16(4):127-133.(in Chinese)
[62] WEISMANN K,HØYER H.Serum alkaline phosphatase and serum zinc levels in the diagnosis and exclusion of zinc deficiency in man[J].The American Journal of Clinical Nutrition,1985,41(6):1214-1219.

[63] QIN S Z,ZHANG L Y,MA F,et al.Dietary zinc and growth,carcass characteristics,immune responses,and serum biochemistry of broilers[J].Animal Production Science,2020,60(6):815-822.

[64] 吕林,陈自宪,罗绪刚.金属硫蛋白在动物体内的功能及其基因表达调控[J].微量元素与健康研究,2010,27(4):58-61. LV L,CHEN Z X,LUO X G.The function and gene expression regulation of metallothionein in animals[J].Studies of Trace Elements and Health,2010,27(4):58-61.(in Chinese)
[65] GRIDER A,BAILEY L B,COUSINS R J.Erythrocyte metallothionein as an index of zinc status in humans[J].Proceedings of the National Academy of Sciences of the United States of America,1990,87(4):1259-1262.

[66] HENNIGAR S R,KELLEY A M,ANDERSON B J,et al.Sensitivity and reliability of zinc transporter and metallothionein gene expression as indicators of zinc status:responses to
ex vivo zinc exposure and habitual zinc intake in humans[J].The British Journal of Nutrition,2020,doi:10.1017/S0007114520002810.
[67] 张军霞.饲粮锌添加水平对蛋鸡产蛋性能、生理生化指标及金属硫蛋白基因表达的影响[D].硕士学位论文.兰州:甘肃农业大学,2004. ZHANG J X.effect of diet supplemented with different zinc levels on production performance,biochemical parameters and expression of metallothionein in laying hen[D].Master's Thesis.Lanzhou:Gansu Agricultural University,2004.(in Chinese)
[68] MIN Y N,LIU F X,QI X,et al.Effects of organic zinc on tibia quality,mineral deposit,and metallothionein expression level of aged hens[J].Poultry Science,2019,98(1):366-372.

[69] WEDEKIND K J,HORTIN A E,BAKER D H.Methodology for assessing zinc bioavailability:efficacy estimates for zinc-methionine,zinc sulfate,and zinc oxide[J].Journal of Animal Science,1992,70(1):178-187.

[70] CAO J,HENRY P R,DAVIS S R,et al.Relative bioavailability of organic zinc sources based on tissue zinc and metallothionein in chicks fed conventional dietary zinc concentrations[J].Animal Feed Science and Technology,2002,101(1/2/3/4):161-170.