[1] 金华,丁家怡,朱庆文.颗粒细胞在未成熟卵体外培养时对未成熟卵母细胞发育的影响[J].齐齐哈尔医学院学报,2006,27(3):309-310.
[2] SHULL M M,DOETSCHMAN T.Transforming growth factor-β1 in reproduction[J].Molecular Reproduction and Development,1994,39(2):239-246.

[3] 郭永红,罗金燕.TGF-β超家族与Smad信号转导研究进展[J].医学综述,2005,11(8):685-688.
[4] DIJKE P T,GOUMANS M J,ITOH F,et al.Regulation of cell proliferation by Smad proteins[J].Journal of Cellular Physiology,2002,191(1):1-16.

[5] GRÖNROOS E,KINGSTON IJ,RAMACHANDRAN A,et al. Transforming Growth Factor β inhibits Bone Morphogenetic Protein-induced transcription through novel phosphorylated Smad1/5-Smad3 complexes[J].Molecular and Cellular Biology,2012,32(14):2904-2916.

[6] 徐梦思.TGFβ-SMAD信号通路对猪颗粒细胞和繁殖性状的作用研究[D].硕士学位论文.石河子:石河子大学,2015:1-20.
[7] 赵瑛,罗玉玉,张文成.TGF-β1/Smads信号通路与eNOS在内皮细胞稳态及动脉粥样硬化发生中的作用[J].武警后勤学院学报:医学版,2013,22(6):560-563.
[8] 张勇,秦娜,于斌.TGF-β/Smads信号转导通路的研究进展[J].广西医科大学学报,2009,26(1):155-157.
[9] 高娜,吴涛.TGF-β在卵巢癌中作用的研究进展[J].大连医科大学学报,2012,34(4):409-413.
[10] LIU S J,DE BOECK M,VAN DAM H,et al.Regulation of the TGF-β pathway by deubiquitinases in cancer[J].The International Journal of Biochemistry & Cell Biology,2016,76:135-145.
[11] BORSUTZKY S,CAZAC B B,ROES J,et al.TGF-β receptor signaling is critical for mucosal IgA responses[J].Journal of Immunology,2004,173(5):3305-3309.

[12] DE CAESTECKER M.The transforming growth factor-β superfamily of receptors[J].Cytokine and Growth Factor Reviews,2004,15(1):1-11.

[13] WANG X F,LIN H Y,NG-EATON E,et al.Expression cloning and characterization of the TGF-β type Ⅲ receptor[J].Cell,1991,67(4):797-805.

[14] 程敏,闻良珍.转化生长因子β及其受体与卵泡发育[J].国外医学:妇产科学分册,2001,28(5):261-264.
[15] PANGAS S A,MATZUK M M.Genetic models for transforming growth factor β superfamily signaling in ovarian follicle development[J].Molecular and Cellular Endocrinology,2004,225(1/2):83-91.
[16] 明佳.Smads蛋白家族与TGF-β的细胞内信号转导[J].国外医学:分子生物学分册,2003,25(2):89-93.
[17] 陈峰,郑敏,陈智.Smad2和Smad3在TGF-β1信号转导中的作用[J].国际流行病学传染病学杂志,2006,33(3):187-189.
[18] 陈骏,钱云良.TGF-β/Smads通路与增生性瘢痕肌成纤维细胞分化[J].中国美容医学杂志,2007,16(7):1000-1003.
[19] TOMIC D,MILLER K P,KENNY H A,et al.Ovarian follicle development requires Smad3[J].Molecular Endocrinology,2004,18(9):2224-2240.

[20] SASSEVILLE M,RITTER L J,NGUYEN T M,et al.Growth differentiation factor 9 signaling requires ERK1/2 activity in mouse granulosa and cumulus cells[J].Journal of Cell Science,2010,123(18):3166-3176.

[21] PANGAS S A,LI X H,ROBERTSON E J,et al.Premature luteinization and cumulus cell defects in ovarian-specific
Smad4 knockout mice[J].Molecular Endocrinology,2006,20(6):1406-1422.

[22] 安新玲,韩金祥,王世立.骨形态发生蛋白的研究进展[J].食品与药品,2009,11(6):69-73.
[23] 王伟.BMP/Smad信号通路对猪卵泡颗粒细胞的影响[D].博士学位论文.南京.南京农业大学,2010:1-13.
[24] 王伟,王少兵,徐银学.BMP/Smad信号通路与哺乳动物卵泡发生[J].遗传,2009,31(3):245-254.
[25] ERICKSON G F,SHIMASAKI S.The spatiotemporal expression pattern of the bone morphogenetic protein family in rat ovary cell types during the estrous cycle[J].Reproductive Biology and Endocrinology,2003,1(1):9.
[26] FENG N P,WANG Z F,ZHANG Z,et al.miR-487b promotes human umbilical vein endothelial cell proliferation,migration,invasion and tube formation through regulating THBS1[J].Neuroscience Letters,2015,591:1-7.
[27] ADAMS J C,LAWLER J.The thrombospondins[J].The International Journal of Biochemistry and Cell Biology,2004,36(6):961-968.

[28] NAGANUMA H,SATOH E,KAWATAKI T,et al.Cell density regulates thrombospondin-1 production in malignant glioma cells[J].Journal of Neuro-Oncology,2003,63(2):147-153.

[29] HUGO C,DANIEL C.Thrombospondin in renal disease[J].Nephron Experimental Nephrology,2009,111(3):e61-e66.
[30] LAWLER J.Thrombospondin-1 as an endogenous inhibitor of angiogenesis and tumor growth[J].Journal of Cellular and Molecular Medicine,2002,6(1):1-12.

[31] ZHOU Z Q,CAO W H,XIE J J,et al.Expression and prognostic significance of THBS1,Cyr61 and CTGF in esophageal squamous cell carcinoma[J].BMC Cancer,2009,9(1):291.
[32] MURPHY-ULLRICH J E,POCZATEK M.Activation of latent TGF-β by thrombospondin-1:mechanisms and physiology[J].Cytokine and Growth Factor Reviews,2000,11(1/2):59-69.
[33] LAWLER J,SUNDAY M,THIBERT V,et al.Thrombospondin-1 is required for normal murine pulmonary homeostasis and its absence causes pneumonia[J].Journal of Clinical Investigation,1998,101(5):982-992.

[34] BAI C,SEN P,HOFMANN K,et al.SKP1 connects cell cycle regulators to the ubiquitin proteolysis machinery through a novel motif,the F-box[J].Cell,1996,86(2):263-274.

[35] FELDMAN R M R,CORRELL C C,KAPLAN K B,et al.A complex of Cdc4p,Skp1p,and Cdc53p/cullin catalyzes ubiquitination of the phosphorylated CDK inhibitor Sic1p[J].Cell,1997,91(2):221-230.

[36] CONNELLY C,HIETER P.Budding yeast
SKP1 encodes an evolutionarily conserved kinetochore protein required for cell cycle progression[J].Cell,1996,86(2):275-285.

[37] 王宇峰.Skp1蛋白在小鼠早期胚胎发育过程中的功能研究[D].硕士学位论文.南京:南京医科大学,2008.