[1] BRUNECKY R,ALAHUHTA M,XU Q,et al.Revealing nature's cellulase diversity:the digestion mechanism of
Caldicellulosiruptor bescii CelA[J].Science,2013,342(6165):1513-1516.

[2] WANG T Y,CHEN H L,LU M Y J,et al.Functional characterization of cellulases identified from the cow rumen fungus
Neocallimastix patriciarum W5 by transcriptomic and secretomic analyses[J].Biotechnology for Biofuels,2011,4:24.
[3] SOLOMON K V,HAITJEMA C H,HENSKE J K,et al.Early-branching gut fungi possess a large,comprehensive array of biomass-degrading enzymes[J].Science,2016,351(6278):1192-1195.

[4] ORPIN C G.Studies on the rumen flagellate
Neocallimastix frontalis[J].Microbiology,1975,91(2):249-262.
[5] ORPIN C G.The occurrence of chitin in the cell walls of the rumen organisms
Neocallimastix frontalis,Piromonas communis and
Sphaeromonas communis[J].Microbiology,1977,99(1):215-218.
[6] BARR D J S.An outline for the reclassification of the Chytridiales,and for a new order,the Spizellomycetales[J].Canadian Journal of Botany,1980,58(22):2380-2394.

[7] FLIEGEROVÁ K,HODROVÁ B,VOIGT K.Classical and molecular approaches as a powerful tool for the characterization of rumen polycentric fungi[J].Folia Microbiologica,2004,49(2):157-164.

[8] GRUNINGER R J,PUNIYA A K,CALLAGHAN T M,et al.Anaerobic fungi (phylum
Neocallimastigomycota):advances in understanding their taxonomy,life cycle,ecology,role and biotechnological potential[J].FEMS Microbiology Ecology,2014,90(1):1-17.

[9] HIBBETT D S,BINDER M,BISCHOFF J F,et al.A higher-level phylogenetic classification of the Fungi[J].Mycological Research,2007,111(5):509-547.

[10] HAITJEMA C H,SOLOMON K V,HENSKE J K,et al.Anaerobic gut fungi:advances in isolation,culture,and cellulolytic enzyme discovery for biofuel production[J].Biotechnology and Bioengineering,2014,111(8):1471-1482.

[11] HO Y W,BARR D J S.Classification of anaerobic gut fungi from herbivores with emphasis on rumen fungi from Malaysia[J].Mycologia,1995,87(5):655-677.

[12] OZKOSE E,THOMAS B J,DAVIES D R,et al.
Cyllamyces aberensis gen.nov.sp.nov.,a new anaerobic gut fungus with branched sporangiophores isolated from cattle[J].Canadian Journal of Botany,2001,79(6):666-673.

[13] HUNGATE R E.A roll tube method for cultivation of strict anaerobes[J].Methods in Microbiology,1969,3:117-132.
[14] CALDWELL D R,BRYANT M P.Medium without rumen fluid for nonselective enumeration and isolation of rumen bacteria[J].Applied Microbiology,1966,14(5):794-801.
[15] LOWE S E,THEODOROU M K,TRINCI A P J,et al.Growth of anaerobic rumen fungi on defined and semi-defined media lacking rumen fluid[J].Microbiology,1985,131(9):2225-2229.

[16] ORPIN C G.Studies on the rumen flagellate
Sphaeromonas communis[J].Microbiology,1976,94(2):270-280.
[17] DORE J,STAHL D A.Phylogeny of anaerobic rumen Chytridiomycetes inferred from small subunit ribosomal RNA sequence comparisons[J].Canadian Journal of Botany,1991,69(9):1964-1971.

[18] SCHOCH C L,SEIFERT K A,HUHNDORF S,et al.Nuclear ribosomal internal transcribed spacer (ITS) region as a universal DNA barcode marker for Fungi[J].Proceedings of the National Academy of Sciences of the United States of America,2012,109(16):6241-6246.

[19] MONCALVO J M,VILGALYS R,REDHEAD S A,et al.One hundred and seventeen clades of euagarics[J].Molecular Phylogenetics and Evolution,2002,23(3):357-400.

[20] WANG X W,LIU X Z,GROENEWALD J Z.Phylogeny of anaerobic fungi (phylum Neocallimastigomycota),with contributions from yak in China[J].Antonie Van Leeuwenhoek,2017,110(1):87-103.

[21] DAGAR S S,KUMAR S,MUDGIL P,et al.D1/D2 domain of large-subunit ribosomal DNA for differentiation of
Orpinomyces spp[J].Applied and Environmental Microbiology,2011,77(18):6722-6725.

[22] TAN H M,CAO L X.Fungal diversity in sheep (
Ovis aries) and cattle (
Bos taurus) feces assessed by comparison of 18S,28S and ITS ribosomal regions[J].Annals of Microbiology,2014,64(3):1423-1427.

[23] BORNEMAN W S,AKIN D E,LJUNGDAHL L G.Fermentation products and plant cell wall-degrading enzymes produced by monocentric and polycentric anaerobic ruminal fungi[J].Applied and Environmental Microbiology,1989,55(5):1066-1073.
[24] LOWE S E,THEODOROU M K,TRINCI A P.Growth and fermentation of an anaerobic rumen fungus on various carbon sources and effect of temperature on development[J].Applied and Environmental Microbiology,1987,53(6):1210-1215.
[25] 李袁飞,孙美洲,成艳芬,等.高效液相色谱法研究瘤胃甲烷菌共存对厌氧真菌代谢产生有机酸特性的影响[J].动物营养学报,2017,29(4):1198-1204.
[26] DEHORITY B A,TIRABASSO P A.Antibiosis between ruminal bacteria and ruminal fungi[J].Applied and Environmental Microbiology,2000,66(7):2921-2927.

[27] JOBLIN K N,MATSUI H,NAYLOR G E,et al.Degradation of fresh ryegrass by methanogenic co-cultures of ruminal fungi grown in the presence or absence of
Fibrobacter succinogenes[J].Current Microbiology,2002,45(1):46-53.

[28] WEI Y Q,YANG H J,LUAN Y,et al.Isolation,identification and fibrolytic characteristics of rumen fungi grown with indigenous methanogen from yaks (
Bos grunniens) grazing on the Qinghai-Tibetan Plateau[J].Journal of Applied Microbiology,2016,120(3):571-587.

[29] WEI Y Q,LONG R J,YANG H,et al.Fiber degradation potential of natural co-cultures of
Neocallimastix frontalis and
Methanobrevibacter ruminantium isolated from yaks (
Bos grunniens) grazing on the Qinghai Tibetan Plateau[J].Anaerobe,2016,39:158-164.
[30] 刁其玉,国春艳.提高粗饲料利用率的途径[J].粮食与饲料工业,2005(10):34-36.
[31] CAO Y C,YANG H J.Effect of roughage fibre content on fibrolytic activities and volatile fatty acid profiles of
Neocallimastix sp.YAK11 isolated from rumen fluids of yak (
Bos grunniens)[J].Animal Feed Science and Technology,2011,170(3/4):284-290.
[32] PAUL S S,DEB S M,PUNIA B S,et al.Fibrolytic potential of anaerobic fungi (
Piromyces sp.) isolated from wild cattle and blue bulls in pure culture and effect of their addition on
in vitro fermentation of wheat straw and methane emission by rumen fluid of buffaloes[J].Journal of the Science of Food and Agriculture,2010,90(7):1218-1226.

[33] NAGPAL R,PUNIYA A K,SEHGAL J P,et al.
In vitro fibrolytic potential of anaerobic rumen fungi from ruminants and non-ruminant herbivores[J].Mycoscience,2011,52(1):31-38.

[34] THAREJA A,PUNIYA A K,GOEL G,et al.
In vitro degradation of wheat straw by anaerobic fungi from small ruminants[J].Archives of Animal Nutrition,2006,60(5):412-417.

[35] LEE S M,GUAN L L,EUN J S,et al.The effect of anaerobic fungal inoculation on the fermentation characteristics of rice straw silages[J].Journal of Applied Microbiology,2015,118(3):565-573.

[36] 王砀砀.瘤胃真菌分离鉴定及其在青贮饲料中应用效果评价[D].硕士学位论文.杨凌:西北农林科技大学,2017:42-48.
[37] LEE S S,HA J K,CHENG K J.Influence of an anaerobic fungal culture administration on
in vivo ruminal fermentation and nutrient digestion[J].Animal Feed Science and Technology,2000,88(3/4):201-217.
[38] SAXENA S,SEHGAL J,PUNIYA A,et al.Effect of administration of rumen fungi on production performance of lactating buffaloes[J].Beneficial Microbes,2010,1(2):183-188.

[39] TRIPATHI V K,SEHGAL J P,PUNIYA A K,et al.Effect of administration of anaerobic fungi isolated from cattle and wild blue bull (
Boselaphus tragocamelus) on growth rate and fibre utilization in buffalo calves[J].Archives of Animal Nutrition,2007,61(5):416-423.

[40] 王砀砀,赵聪聪,郭奇奇,等.瘤胃真菌分离鉴定及酶学特性[J].农业生物技术学报,2017,25(10):1668-1681.
[41] CHEN Y C,CHIANG Y C,HSU F Y,et al.Structural modeling and further improvement in pH stability and activity of a highly-active xylanase from an uncultured rumen fungus[J].Bioresource Technology,2012,123:125-134.
[42] 曹阳春.牦牛瘤胃厌氧真菌多样性与饲料细胞壁降解酯酶特性研究[D].博士学位论文.北京:中国农业大学,2012:70-75.
[43] RANGANATHAN A,SMITH O P,YOUSSEF N H,et al.Utilizing anaerobic fungi for two-stage sugar extraction and biofuel production from lignocellulosic biomass[J].Frontiers in Microbiology,2017,8:635.
[44] CHUNG D H,CHA M S,GUSS A M,et al.Direct conversion of plant biomass to ethanol by engineered
Caldicellulosiruptor bescii[J].Proceedings of the National Academy of Sciences of the United States of America,2014,111(24):8931-8936.

[45] JIN M J,GUNAWAN C,BALAN V,et al.Consolid