[1] ZHU H,YANG T,XING Y X,et al.Advances in nutritional function of extracellular polysaccharides from lactic acid bacteria[J].Revista Cientifica-Facultad De Ciencias Veterinarias,2020,30(1):345-353.
[2] LEBLANC J G,LEVIT R,SAVOY DE GIORI G,et al.Application of vitamin-producing lactic acid bacteria to treat intestinal inflammatory diseases[J].Applied Microbiology and Biotechnology,2020,104(8):3331-3337.
[3] KIM K W,KANG S S,WOO S J,et al.Lipoteichoic acid of probiotic
Lactobacillus plantarum attenuates poly Ⅰ:C-induced IL-8 production in porcine intestinal epithelial cells[J].Frontiers in Microbiology,2017,8:1827.
[4] ZEIDAN A A,POULSEN V K,JANZEN T,et al.Polysaccharide production by lactic acid bacteria:from genes to industrial applications[J].FEMS Microbiology Reviews,2017,41(Suppl.1):S168-S200.
[5] XIU L,SHENG S X,HU Z P,et al.Exopolysaccharides from
Lactobacillus kiferi as adjuvant enhanced the immuno-protective against
Staphylococcus aureus infection[J].International Journal of Biological Macromolecules,2020,161:10-23.
[6] DI W,ZHANG L W,WANG S M,et al.Physicochemical characterization and antitumour activity of exopolysaccharides produced by
Lactobacillus casei SB27 from yak milk[J].Carbohydrate Polymers,2017,171:307-315.
[7] SANALIBABA P,CAKMAK G A.Exopolysaccharides production by lactic acid bacteria[J].Applied Microbiology:Open Access,2016,2(2):1000115.
[8] LYNCH K M,ZANNINI E,COFFEY A,et al.Lactic acid bacteria exopolysaccharides in foods and beverages:isolation,properties,characterization,and health benefits[J].Annual Review of Food Science and Technology,2018,9:155-176.
[9] WANG K,LI W,RUI X,et al.Characterization of a novel exopolysaccharide with antitumor activity from
Lactobacillus plantarum 70810[J].International Journal of Biological Macromolecules,2014,63:133-139.
[10] SIRICHOKCHATCHAWAN W,TEMEEYASEN G,NILUBOL D,et al.Protective effects of cell-free supernatant and live lactic acid bacteria isolated from thai pigs against a pandemic strain of porcine epidemic diarrhea virus[J].Probiotics and Antimicrobial Proteins,2018,10(2):383-390.
[11] LIU Z Q,ZHANG Z H,QIU L,et al.Characterization and bioactivities of the exopolysaccharide from a probiotic strain of
Lactobacillus plantarum WLPL04[J].Journal of Dairy Science,2017,100(9):6895-6905.
[12] AYYASH M,ABU-JDAYIL B,ITSARANUWAT P,et al.Exopolysaccharide produced by the potential probiotic
Lactococcus garvieae C47:structural characteristics,rheological properties,bioactivities and impact on fermented camel milk[J].Food Chemistry,2020,333:127418.
[13] ANGELIN J,KAVITHA M.Exopolysaccharides from probiotic bacteria and their health potential[J].International Journal of Biological Macromolecules,2020,162:853-865.
[14] XU Y M,CUI Y L,YUE F F,et al.Exopolysaccharides produced by lactic acid bacteria and
Bifidobacteria:structures,physiochemical functions and applications in the food industry[J].Food Hydrocolloids,2019,94:475-499.
[15] LI W,TANG W Z,JI J,et al.Characterization of a novel polysaccharide with anti-colon cancer activity from
Lactobacillus helveticus MB2-1[J].Carbohydrate Research,2015,411:6-14.
[16] LI W,JI J,CHEN X H,et al.Structural elucidation and antioxidant activities of exopolysaccharides from
Lactobacillus helveticus MB2-1[J].Carbohydrate Polymers,2014,102:351-359.
[17] MAKINO S,SATO A,GOTO A,et al.Enhanced natural killer cell activation by exopolysaccharides derived from yogurt fermented with
Lactobacillus delbrueckii ssp.
bulgaricus OLL1073R-1[J].Journal of Dairy Science,2016,99(2):915-923.
[18] ZHOU K,ZENG Y T,YANG M L,et al.Production,purification and structural study of an exopolysaccharide from
Lactobacillus plantarum BC-25[J].Carbohydrate Polymers,2016,144:205-214.
[19] AI L Z,GUO Q B,DING H H,et al.Structure characterization of exopolysaccharides from
Lactobacillus casei LC2W from skim milk[J].Food Hydrocolloids,2016,56:134-143.
[20] GÓRSKA-FRACZEK S,SANDSTRÖM C,KENNE L,et al.The structure and immunoreactivity of exopolysaccharide isolated from
Lactobacillus johnsonii strain 151[J].Carbohydrate Research,2013,378:148-153.
[21] FONTANA C,LI S Y,YANG Z N,et al.Structural studies of the exopolysaccharide from
Lactobacillus plantarum C88 using NMR spectroscopy and the program CASPER[J].Carbohydrate Research,2015,402:87-94.
[22] GÓRSKA-FRACZEK S,SANDSTRÖM C,KENNE L,et al.Structural studies of the exopolysaccharide consisting of a nonasaccharide repeating unit isolated from
Lactobacillus rhamnosus KL37B[J].Carbohydrate Research,2011,346(18):2926-2932.
[23] SÄWÉN E,HUTTUNEN E,ZHANG X,et al.Structural analysis of the exopolysaccharide produced by
Streptococcus thermophilus ST1 solely by NMR spectroscopy[J].Journal of Biomolecular NMR,2010,47(2):125-134.
[24] MARSHALL V M,DUNN H,ELVIN M,et al.Structural characterisation of the exopolysaccharide produced by
Streptococcus thermophilus EU20[J].Carbohydrate Research,2001,331(4):413-422.
[25] SHIOMI M,SASAKI K,MUROFUSHI M,et al.Antitumor activity in mice of orally administered polysaccharide from kefir grain[J].Japanese Journal of Medical Science and Biology,1982,35(2):75-80.
[26] LI L,JIANG Y J,YANG X Y,et al.Immunoregulatory effects on Caco-2 cells and mice of exopolysaccharides isolated from
Lactobacillus acidophilus NCFM[J].Food & Function,2014,5(12):3261-3268.
[27] EL-DEEB N M,YASSIN A M,AL-MADBOLY L A,et al.A novel purified
Lactobacillus acidophilus 20079 exopolysaccharide,LA-EPS-20079,molecularly regulates both apoptotic and NF-κB inflammatory pathways in human colon cancer[J].Microbial Cell Factories,2018,17(1):29.
[28] SUN N X,LIU H P,LIU S J,et al.Purification,preliminary structure and antitumor activity of exopolysaccharide produced by
Streptococcus thermophilus CH9[J].Molecules,2018,23(11):2898.
[29] 胡盼盼.乳酸菌胞外多糖诱导结肠癌HT-29细胞凋亡的机制研究[D].硕士学位论文.哈尔滨:哈尔滨工业大学,2015. HU P P.The mechanism of exopolysaccharide-induced apoptosis in HT-29 cancer cell[D].Master's Thesis.Harbin:Harbin Institute of Technology,2015.(in Chinese)
[30] SCHIEBER M,CHANDEL N S.ROS function in redox signaling and oxidative stress[J].Current Biology,2014,24(10):R453-R462.
[31] CAO P,ZHANG Y,HUANG Z,et al.The preventative effects of procyanidin on binge ethanol-induced lipid accumulation and ROS overproduction via the promotion of hepatic autophagy[J].Molecular Nutrition & Food Research,2019,63(18):1801255.
[32] RAGURAMAN V,L S A,J J,et al.Sulfated polysaccharide from
Sargassum tenerrimum attenuates oxidative stress induced reactive oxygen species production in
in vitro and in zebrafish model[J].Carbohydrate Polymers,2019,203:441-449.
[33] KRUK J,ABOUL-ENEIN H,KŁADNA A,et al.Oxidative stress in biological systems and its relation with pathophysiological functions:the effect of physical activity on cellular redox homeostasis[J].Free Radical Research,2019,53(5):497-521.
[34] LI S Y,ZHAO Y J,ZHANG L,et al.Antioxidant activity of
Lactobacillus plantarum strains isolated from traditional Chinese fermented foods[J].Food Chemistry,2012,135(3):1914-1919.
[35] MIN W H,FANG X B,WU T,et al.Characterization and antioxidant activity of an acidic exopolysaccharide from
Lactobacillus plantarum JLAU103[J].Journal of Bioscience and Bioengineering,2019,127(6):758-766.
[36] TANG W Z,DONG M S,WANG W L,et al.Structural characterization and antioxidant property of released exopolysaccharides from
Lactobacillus delbrueckii ssp.
bulgaricus SRFM-1[J].Carbohydrate Polymers,2017,173:654-664.
[37] WANG X,SHAO C G,LIU L,et al.Optimization,partial characterization and antioxidant activity of an exopolysaccharide from
Lactobacillus plantarum KX041[J].International Journal of Biological Macromolecules,2017,103:1173-1184.
[38] 刘煜珺,张雨晴,高原,等.乳杆菌胞外多糖抗氧化活性研究[J].中国食品学报,2019,19(6):21-35. LIU Y J,ZHANG Y Q,GAO Y,et al.Studies on the antioxidative activity of exopolysaccharide from
Lactobacillus strains[J].Journal of Chinese Institute of Food Science and Technology,2019,19(06):21-35.(in Chinese)
[39] 白丽娟.马奶酒中产胞外多糖瑞士乳杆菌的筛选及多糖的结构和抗氧化活性研究[D].博士学位论文.沈阳:沈阳农业大学,2017. BAI L J.Screening exopolysaccharide-producing
L. helveticus and structure,antioxidant activity of exopolysaccharides[D].Ph.D. Thesis.Shenyang:Shenyang Agricultural University,2017.(in Chinese)
[40] OLEKSY-SOBCZAK M,KLEWICKA E.Optimization of media composition to maximize the yield of exopolysaccharides production by
Lactobacillus rhamnosus strains[J].Probiotics and Antimicrobial Proteins,2020,12(2):774-783.
[41] XIAO L Y,LI Y Y,TIAN J J,et al.Influences of drying methods on the structural,physicochemical and antioxidant properties of exopolysaccharide from
Lactobacillus helveticus MB2-1[J].International Journal of Biological Macromolecules,2020,157:220-231.
[42] RIAZ RAJOKA M S,JIN M L,ZHAO H B,et al.Functional characterization and biotechnological potential of exopolysaccharide produced by
Lactobacillus rhamnosus strains isolated from human breast milk[J].LWT,2018,89:638-647.
[43] ZHANG J,ZHAO X,JIANG Y Y,et al.Antioxidant status and gut microbiota change in an aging mouse model as influenced by exopolysaccharide produced by
Lactobacillus plantarum YW11 isolated from Tibetan kefir[J].Journal of Dairy Science,2017,100(8):6025-6041.
[44] TRABELSI I,KTARI N,BEN SLIMA S,et al.Evaluation of dermal wound healing activity and
in vitro antibacterial and antioxidant activities of a new exopolysaccharide produced by
Lactobacillus sp. Ca6[J].International Journal of Biological Macromolecules,2017,103:194-201.
[45] ABEDFAR A,ABBASZADEH S,HOSSEININEZHAD M,et al.Physicochemical and biological characterization of the EPS produced by
L. acidophilus isolated from rice bran sourdough[J].LWT,2020,127:109373.
[46] WANG K,LI W,RUI X,et al.Structural characterization and bioactivity of released exopolysaccharides from
Lactobacillus plantarum 70810[J].International Journal of Biological Macromolecules,2014,67:71-78.
[47] LI S Q,SHAH N P.Antioxidant and antibacterial activities of sulphated polysaccharides from
Pleurotus eryngii and
Streptococcus thermophilus ASCC 1275[J].Food Chemistry,2014,165:262-270.
[48] MAO J W,YIN J,GE Q,et al.
In vitro antioxidant activities of polysaccharides extracted from
Moso Bamboo-Leaf[J].International Journal of Biological Macromolecules,2013,55:1-5.
[49] KANMANI P,ALBARRACIN L,KOBAYASHI H,et al.Exopolysaccharides from
Lactobacillus delbrueckii OLL1073R-1 modulate innate antiviral immune response in porcine intestinal epithelial cells[J].Molecular Immunology,2018,93:253-265.
[50] KIM K,LEE G,THANH H D,et al.Exopolysaccharide from
Lactobacillus plantarum LRCC5310 offers protection against rotavirus-induced diarrhea and regulates inflammatory response[J].Journal of Dairy Science,2018,101(7):5702-5712.
[51] KANMANI P,ALBARRACIN L,KOBAYASHI H,et al.Genomic characterization of
Lactobacillus delbrueckii TUA4408L and evaluation of the antiviral activities of its extracellular polysaccharides in porcine intestinal epithelial cells[J].Frontiers in Immunology,2018,9:2178.
[52] 王莹莹.嗜酸乳杆菌胞外多糖对TGEV感染ST细胞TLR3及部分细胞因子表达的影响[D].硕士学位论文.哈尔滨:东北农业大学,2015. WANG Y Y.The expression of TLR3 and some cytokines in ST cells induced by TGEV infection and
Lactobacillus acidophilus exopolysaccharides pretreatment[D].Master's Thesis.Harbin:Northeast Agricultural University,2015.(in Chinese)
[53] ZHU Y T,WANG X J,PAN W S,et al.Exopolysaccharides produced by yogurt-texture improving
Lactobacillus plantarum RS20D and the immunoregulatory activity[J].International Journal of Biological Macromolecules,2019,121:342-349.
[54] 李琪,杜佳峰,高洁,等.副干酪乳杆菌VL8胞外多糖对巨噬细胞RAW264.7免疫激活作用研究[J].食品研究与开发,2020,41(3):208-213. LI Q,DU J F,GAI J,et al.Immune activation of macrophage RAW264.7 by
Lactobacillus paracasei VL8 exopolysaccharides[J].Food Research and Development,2020,41(3):208-213.(in Chinese)
[55] BALZARETTI S,TAVERNITI V,GUGLIELMETTI S,et al.A novel rhamnose-rich hetero-exopolysaccharide isolated from
Lactobacillus paracasei DG activates THP-1 human monocytic cells[J].Applied and Environmental Microbiology,2017,83(3):e02702-16.
[56] ZHOU Y,CUI Y H,QU X J.Exopolysaccharides of lactic acid bacteria:structure,bioactivity and associations:a review[J].Carbohydrate Polymers,2019,207:317-332.
[57] KORCZ E,KERÉNYI Z,VARGA L.Dietary fibers,prebiotics,and exopolysaccharides produced by lactic acid bacteria:potential health benefits with special regard to cholesterol-lowering effects[J].Food & Function,2018,9(6):3057-3068.
[58] CHAISUWAN W,JANTANASAKULWONG K,WANGTUEAI S,et al.Microbial exopolysaccharides for immune enhancement:fermentation,modifications and bioactivities[J].Food Bioscience,2020,35:100564.
[59] TANG C,DING R X,SUN J,et al.The impacts of natural polysaccharides on intestinal microbiota and immune responses-a review[J].Food & Function,2019,10(5):2290-2312.
[60] 童良琴,曲亚军,陈敏.乳酸菌胞外多糖的研究进展[J].中国生物工程杂志,2015,35(11):85-91. TONG L Q,QU Y J,CHEN M.Research advance on exopolysaccharides synthesized by lactic acid bacteria[J].China Biotechnology,2015,35(11):85-91.(in Chinese)
[61] LI S J,HUANG R H,SHAH N P,et al.Antioxidant and antibacterial activities of exopolysaccharides from
Bifidobacterium bifidum WBIN03 and
Lactobacillus plantarum R315[J].Journal of Dairy Science,2014,97(12):7334-7343.
[62] JEONG D,KIM D H,KANG I B,et al.Characterization and antibacterial activity of a novel exopolysaccharide produced by
Lactobacillus kefiranofaciens DN1 isolated from kefir[J].Food Control,2017,78:436-442.
[63] ALLONSIUS C N,VAN DEN BROEK M F L,DE BOECK I,et al.Interplay between
Lactobacillus rhamnosus GG and
Candida and the involvement of exopolysaccharides[J].Microbial Biotechnology,2017,10(6):1753-1763.
[64] WANG J,ZHAO X,YANG Y W,et al.Characterization and bioactivities of an exopolysaccharide produced by
Lactobacillus plantarum YW32[J].International Journal of Biological Macromolecules,2015,74:119-126.
[65] 邵丽.产胞外多糖乳杆菌的筛选及其多糖的分离、结构和生物活性研究[D].博士学位论文.无锡:江南大学,2015. SHAO L.Screening of exopolysaccharide-producing lactobacilli and separation,structure and bioactivities of exopolysaccharide[D].Ph.D. Thesis.Wuxi:Jiangnan University,2015.(in Chinese)
[66] 李胜杰.两歧双歧杆菌WBIN03胞外多糖的抗菌/抗氧化活性及其对小鼠免疫和肠道菌群影响的研究[D].硕士学位论文.南昌:南昌大学,2014. LI S J.Research on the antioxidant and antibacterial active of exopolysaccharides from
Bifidobacterium bifidum WBIN03 and its benefical effects on the immunity and inteatinal microbiota in mice[D].Master's Thesis.Nanchang:Nanchang University,2014.(in Chinese)
[67] KANMANI P,SUGANYA K,KUMAR R S,et al.Synthesis and functional characterization of antibiofilm exopolysaccharide produced by
Enterococcus faecium MC13 isolated from the gut of fish[J].Applied Biochemistry and Biotechnology,2013,169(3):1001-1015.
[68] DINIĆ M,PECIKOZA U,DJOKIĆ J,et al.Exopolysaccharide produced by probiotic strain
Lactobacillus paraplantarum BGCG11 reduces inflammatory hyperalgesia in rats[J].Frontiers in Pharmacology,2018,9:1.
[69] FOOLADI T,SOUDI M R,ALIMADADI N,et al.Bioactive exopolysaccharide from
Neopestalotiopsis sp. strain SKE15:production,characterization and optimization[J].International Journal of Biological Macromolecules,2019,129:127-139.
[70] AI L Z,ZHANG H,GUO B H,et al.Preparation,partial characterization and bioactivity of exopolysaccharides from
Lactobacillus casei LC2W[J].Carbohydrate Polymers,2008,74(3):353-357.
[71] MAEDA H,ZHU X,SUZUKI S,et al.Structural characterization and biological activities of an exopolysaccharide kefiran produced by
Lactobacillus kefiranofaciens WT-2B[J].Journal of Agricultural and Food Chemistry,2004,52(17):5533-5538.
[72] SASIKUMAR K,KOZHUMMAL VAIKKATH D,DEVENDRA L,et al.An exopolysaccharide (EPS) from a
Lactobacillus plantarum BR2 with potential benefits for making functional foods[J].Bioresource Technology,2017,241:1152-1156.
[73] ZHANG H Y,PENG J X,LI X,et al.A nano-bio interfacial protein corona on silica nanoparticle[J].Colloids and Surfaces B:Biointerfaces,2018,167:220-228.
[74] HAN X,YANG Z,JING X P,et al.Improvement of the texture of yogurt by use of exopolysaccharide producing lactic acid bacteria[J].Biomed Research International,2016,2016:7945675.
[75] LLUIS-ARROYO D,FLORES-NÁJERA A,CRUZ-GUERRERO A,et al.Effect of an exopolysaccharide-producing strain of
Streptococcus thermophilus on the yield and texture of mexican manchego-type cheese[J].International Journal of Food Properties,2014,17(8):1680-1693.
[76] DERTLI E,MAYER M J,COLQUHOUN I J,et al.EpsA is an essential gene in exopolysaccharide production in
Lactobacillus johnsonii FI9785[J].Microbial Biotechnology,2016,9(4):496-501.
[77] LYNCH K M,COFFEY A,ARENDT E K.Exopolysaccharide producing lactic acid bacteria:their techno-functional role and potential application in gluten-free bread products[J].Food Research International,2018,110:52-61.
[78] TOUTAIN P L,FERRAN A A,BOUSQUET-MELOU A,et al.Veterinary medicine needs new green antimicrobial drugs[J].Frontiers in Microbiology,2016,7:1196.
[79] NEUMAN H,FORSYTHE P,UZAN A,et al.Antibiotics in early life:dysbiosis and the damage done[J].FEMS Microbiology Reviews,2018,42(4):489-499.
[80] WANG J,JI H F,WANG S X,et al.Probiotic
Lactobacillus plantarum promotes intestinal barrier function by strengthening the epithelium and modulating gut microbiota[J].Frontiers in Microbiology,2018,9:1953.
[81] AZAD M A K,SARKER M,LI T J,et al.Probiotic species in the modulation of gut microbiota:an overview[J].Biomed Research International,2018,2018:9478630.
[82] CAGGIANIELLO G,KLEEREBEZEM M,SPANO G.Exopolysaccharides produced by lactic acid bacteria:from health-promoting benefits to stress tolerance mechanisms[J].Applied Microbiology and Biotechnology,2016,100(9):3877-3886.
[83] FANNING S,HALL L J,CRONIN M,et al.Bifidobacterial surface-exopolysaccharide facilitates commensal-host interaction through immune modulation and pathogen protection[J].Proceedings of the National Academy of Sciences of the United States of America,2012,109(6):2108-2113.
[84] YANG Y,GALLE S,LE M H,et al.Feed fermentation with reuteran- and levan-producing
Lactobacillus reuteri reduces colonization of weanling pigs by enterotoxigenic
Escherichia coli[J].Applied and Environmental Microbiology,2015,81(17):5743-5752.
[85] ASHFAQ I,AMJAD H,AHMAD W,et al.Growth inhibition of common enteric pathogens in the intestine of broilers by microbially produced dextran and levan exopolysaccharides[J].Current Microbiology,2020,77(9):2128-2136.
[86] RAJOKA M S R,MEHWISH H M,HAYAT H F,et al.Characterization,the antioxidant and antimicrobial activity of exopolysaccharide isolated from poultry origin
Lactobacilli[J].Probiotics and Antimicrobial Proteins,2019,11(4):1132-1142.
