[1] ADIBI S A,MERCER D W.Protein digestion in human intestine as reflected in luminal,mucosal,and plasma amino acid concentrations after meals[J].Journal of Clinical Investigation,1973,52(7):1586-1594.

[2] DANIEL H.Molecular and integrative physiology of intestinal peptide transport[J].Annual Review of Physiology,2004,66:361-384.
[3] VERRI T,BARCA A,PISANI P,et al.Di- and tripeptide transport in vertebrates:the contribution of teleost fish models[J].Journal of Comparative Physiology B,2017,187(3):395-462.

[4] NEWSTEAD S.Recent advances in understanding proton coupled peptide transport via the POT family[J].Current Opinion in Structural Biology,2017,45:17-24.
[5] LU H,KLAASSEN C.Tissue distribution and thyroid hormone regulation of
Pept1 and
Pept2 mRNA in rodents[J].Peptides,2006,27(4):850-857.

[6] ZWARYCZ B,WONG E A.Expression of the peptide transporters
PepT1,
PepT2,and
PHT1 in the embryonic and posthatch chick[J].Poultry Science,2013,92(5):1314-1321.

[7] PAN Y X,WONG E A,BLOOMQUIST J R,et al.Expression of a cloned ovine gastrointestinal peptide transporter (oPepT1) in
Xenopus oocytes induces uptake of oligopeptides in vitro[J].The Journal of Nutrition,2001,131(4):1264-1270.

[8] FEI Y J,KANAI Y,NUSSBERGER S,et al.Expression cloning of a mammalian proton-coupled oligopeptide transporter[J].Nature,1994,368(6471):563-566.

[9] HART H R,EVANS A N,GELSLEICHTER J,et al.Molecular identification and functional characteristics of peptide transporters in the Bonnethead shark (
Sphyrna tiburo)[J].Journal of Comparative Physiology B,2016,186(7):855-866.

[10] VACCA F,BARCA A,GOMES A S,et al.The peptide transporter 1a of the zebrafish
Danio rerio,an emerging model in nutrigenomics and nutrition research:molecular characterization,functional properties,and expression analysis[J].Genes and Nutrition,2019,14:33.
[11] SMITH D E,CLÉMENÇON B,HEDIGER M A.Proton-coupled oligopeptide transporter family SLC15:physiological,pharmacological and pathological implications[J].Molecular Aspects of Medicine,2013,34(2/3):323-336.
[12] OPPERMANN H,HEINRICH M,BIRKEMEYER C,et al.The proton-coupled oligopeptide transporters PEPT2,PHT1 and PHT2 mediate the uptake of carnosine in glioblastoma cells[J].Amino Acids,2019,51(7):999-1008.

[13] LIANG R,FEI Y J,PRASAD P D,et al.Human intestinal H
+/peptide cotransporter:cloning,functional expression,and chromosomal localization[J].Journal of Biological Chemistry,1995,270(12):6456-6463.

[14] SAITO H,MOTOHASHI H,MUKAI M,et al.Cloning and characterization of a pH-sensing regulatory factor that modulates transport activity of the human H
+/peptide cotransporter,PEPT1[J].Biochemical and Biophysical Research Communications,1997,237(3):577-582.

[15] BOLL M,MARKOVICH D,WEBER W M,et al.Expression cloning of a cDNA from rabbit small intestine related to proton-coupled transport of peptides,βlactam antibiotics and ACE-inhibitors[J].Pflügers Archiv,1994,429(1):146-149.

[16] SAITO H,OKUDA M,TERADA T,et al.Cloning and characterization of a rat H
+/peptide cotransporter mediating absorption of beta-lactam antibiotics in the intestine and kidney[J].Journal of Pharmacology and Experimental Therapeutics,1995,275(3):1631-1637.
[17] FEI Y J,SUGAWARA M,LIU J C,et al.cDNA structure,genomic organization,and promoter analysis of the mouse intestinal peptide transporter PEPT1[J].Biochimica et Biophysica Acta (BBA):Gene Structure and Expression,2000,1492(1):145-154.

[18] KLANG J E,BURNWORTH L A,PAN Y X,et al.Functional characterization of a cloned pig intestinal peptide transporter (pPepT1)[J].Journal of Animal Science,2005,83(1):172-181.

[19] FRAZIER S,AJIBOYE K,OLDS A,et al.Functional characterization of the chicken peptide transporter 1(
pept1,
slc15
a1) gene[J].Animal Biotechnology,2008,19(4):201-210.

[20] ZIMIN A V,DELCHER A L,FLOREA L,et al.A whole-genome assembly of the domestic cow,
Bos taurus[J].Genome Biology,2009,10(4):R42.
[21] XU Q B,LIU Z X,LIU H Y,et al.Functional characterization of oligopeptide transporter 1 of dairy cows[J].Journal of Animal Science and Biotechnology,2018,9(1):7.
[22] CHEN H,PAN Y X,WONG E A,et al.Characterization and regulation of a cloned ovine gastrointestinal peptide transporter (
oPepT1) expressed in a mammalian cell line[J].The Journal of Nutrition,2002,132(1):38-42.

[23] WANG H C,SHI F Y,HOU M J,et al.Cloning of oligopeptide transport carrier PepT1 and comparative analysis of PepT1 messenger ribonucleic acid expression in response to dietary nitrogen levels in yak (
Bos grunniens) and indigenous cattle (
Bos taurus) on the
Qinghai-Tibetan plateau[J].Journal of Animal Science,2016,94(8):3431-3440.

[24] AMBERG J J,MYR C,KAMISAKA Y,et al.Expression of the oligopeptide transporter,
PepT1,in larval Atlantic cod (
Gadus morhua)[J].Comparative Biochemistry and Physiology Part B:Biochemistry and Molecular Biology,2008,150(2):177-182.

[25] GOMES A S,VACCA F,CINQUETTI R,et al.Identification and characterization of the Atlantic salmon peptide transporter 1a[J].American Journal of Physiology:Cell Physiology,2020,318(1):C191-C204.
[26] DANIEL H,KOTTRA G.The proton oligopeptide cotransporter family SLC15 in physiology and pharmacology[J].Pflügers Archiv,2004,447(5):610-618.

[27] SHIMIZU Y,MASUDA S,NISHIHARA K,et al.Increased protein level of PEPT1 intestinal H
+-peptide cotransporter upregulates absorption of glycylsarcosine and ceftibuten in 5/6 nephrectomized rats[J].Gastrointestinal and Liver Physiology,2004,288(4):G664-G670.
[28] ALGHAMDI O A,KING N,ANDRONICOS N M,et al.Molecular changes to the rat renal cotransporters PEPT1 and PEPT2 due to ageing[J].Molecular and Cellular Biochemistry,2019,452(1/2):71-82.
[29] 张雨,丰凯,汤方.杨小舟蛾
MtroOBP1基因的克隆、序列分析及表达[J].南京林业大学学报(自然科学版),2020,44(5):219-225. ZHANG Y,FENG K,TANG F.The cloning,sequence analysis and expression of
MtroOBP1 gene in
Micromelalopha troglodyte (Graeser)[J].Journal of Nanjing Forestry University (Natural Sciences Edition),2020,44(5):219-225.(in Chinese)
[30] LIVAK K J,SCHMITTGEN T D.Analysis of relative gene expression data using real-time quantitative PCR[J].Methods,2002,25(4):402-408.
[31] DANIEL H,SPANIER B,KOTTRA G,et al.From bacteria to man:archaic proton-dependent peptide transporters at work[J].Physiology,2006,21(2):93-102.

[32] NEWSTEAD S.Towards a structural understanding of drug and peptide transport within the proton-dependent oligopeptide transporter (POT) family[J].Biochemical Society Transactions,2011,39(5):1353-1358.

[33] JENKINS N,PAREKH R B,JAMES D C.Getting the glycosylation right:implications for the biotechnology industry[J].Nature Biotechnology,1996,14(8):975-981.

[34] CHAN T,LU X X,SHAMS T,et al.The role of N-glycosylation in maintaining the transporter activity and expression of human oligopeptide transporter[J].Molecular Pharmaceutics,2016,13(10):3449-3456.

[35] WANG C H,ZHAO F Q,LIU J X,et al.Short communication:the essential role of N-glycosylation in the transport activity of bovine peptide transporter 2[J].Journal of Dairy Science,2020,103(7):6679-6683.

[36] HIJO A H T,COUTINHO C P,ALBA-LOUREIRO T C,et al.High fat diet modulates the protein content of nutrient transporters in the small intestine of mice:possible involvement of PKA and PKC activity[J].Heliyon,2019,5(10):e02611.
[37] BRANDSCH M,MIYAMOTO Y,GANAPATHY V,et al.Expression and protein kinase C-dependent regulation of peptide/H
+ co-transport system in the Caco-2 human colon carcinoma cell line[J].Biochemical Journal,1994,299(1):253-260.

[38] WENZEL U,DIEHL D,HERGET M,et al.Regulation of the high-affinity H
+/peptide cotransporter in renal LLC-PK1 cells[J].Journal of Cellular Physiology,1999,178(3):341-348.

[39] DANIEL H,ZIETEK T.Taste and move:glucose and peptide transporters in the gastrointestinal tract[J].Experimental Physiology,2015,100(12):1441-1450.

[40] HUANG Q S,DELGADO J M V,PINOARGOTE O D S,et al.Molecular evolution of the Slc15 family and its response to waterborne copper and mercury exposure in tilapia[J].Aquatic Toxicology,2015,163:140-147.
[41] 卢艳娟.日粮能量蛋白水平对滩羊主要营养素转运载体mRNA表达的影响[D].硕士学位论文.杨凌:西北农林科技大学,2017. LU Y J.Effects of diets with different energy and protein levels on mrna expression of nutrient transporters in
Tan sheep[D].Master's Thesis.Yangling:Northwest Agriculture & Forestry University,2007.(in Chinese)