[1] 王虎成, 龙瑞军.青藏高原牦牛生产现状与展望[C]//2009中国草原发展论坛论文集.合肥:中国草学会, 2009:118-121. WANG H C, LONG R J.Present situation and prospect of yak production in
Qinghai Tibet Plateau[C]//Proceedings of 2009 Chinese Grassland Development Forum.Hefei:Chinese Grassland Society, 2009:181-121.(in Chinese)
[2] 巴桑措姆.牦牛养殖发展现状及对策[J].畜牧兽医科学(电子版), 2020(9):70-71. BA S C M.Status and countermeasures of yak breeding[J].Graziery Veterinary Sciences (Electronic Version), 2020(9):70-71.(in Chinese)
[3] 骆正杰, 马进寿, 保广才, 等.青海省牦牛种业发展现状、存在问题及应对策略[J].中国畜牧杂志, 2021, 57(2):231-234. LUO Z J, MA J S, BAO G C, et al.Development status, existing problems and countermeasures of yak seed industry in
Qinghai Province[J].Chinese Journal of Animal Science, 2021, 57(2):231-234.(in Chinese)
[4] 吴发莉, 王之盛, 杨勤, 等.甘南碌曲和合作地区冬夏季高寒天然牧草生产特性、营养成分和饲用价值分析[J].草业学报, 2014, 23(4):31-38. WU F L, WANG Z S, YANG Q, et al.Analysis of growth characteristics, nutritional components and feeding values of native forage grass from the high-cold steppes in the
Luqu and
Hezuo region of
Gannan in summer and winter[J].Acta Prataculturae Sinica, 2014, 23(4):31-38.(in Chinese)
[5] 王威, 张建勋, 康坤, 等.冷季补饲精料对牦牛繁殖性能和生长性能的影响[J].中国畜牧杂志, 2013, 49(7):78-80. WANG W, ZHANG J X, KANG K, et al.Effects of cold season supplementary feeding concentrate on reproductive performance and growth performance of yak[J].Chinese Journal of Animal Science, 2013, 49(7):78-80.(in Chinese)
[6] MCCUE M D.Starvation physiology:reviewing the different strategies animals use to survive a common challenge[J].Comparative Biochemistry and Physiology-Part A:Molecular & Integrative Physiology, 2010, 156(1):1-18.

[7] SINGH S P, HÄUSSLER S, HEINZ J F L, et al.Lactation driven dynamics of adiponectin supply from different fat depots to circulation in cows[J].Domestic Animal Endocrinology, 2014, 47:35-46.
[8] BRADY L J, ROMSOS D R, BRADY P S, et al.The effects of fasting on body composition, glucose turnover, enzymes and metabolites in the chicken[J].The Journal of Nutrition, 1978, 108(4):648-657.

[9] DAVIS K B, GAYLORD T G.Effect of fasting on body composition and responses to stress in sunshine bass[J].Comparative Biochemistry and Physiology-Part A:Molecular & Integrative Physiology, 2011, 158(1):30-36.

[10] DEDKOVA E N, BLATTER L A.Role of β-hydroxybutyrate, its polymer poly-β-hydroxybutyrate and inorganic polyphosphate in mammalian health and disease[J].Frontiers in Physiology, 2014, 5:260.
[11] HAHN P, TALLER M.Ketone formation in the intestinal mucosa of infant rats[J].Life Sciences, 1987, 41(12):1525-1528.

[12] HEGARDT F G.Mitochondrial 3-hydroxy-3-methylglutaryl-CoA synthase:a control enzyme in ketogenesis[J].Biochemical Journal, 1999, 338(Pt 3):569-582.
[13] THUMELIN S, FORESTIER M, GIRARD J, et al.Developmental changes in mitochondrial 3-hydroxy-3-methylglutaryl-CoA synthase gene expression in rat liver, intestine and kidney[J].Biochemical Journal, 1993, 292(Pt 2):493-496.
[14] ROJAS-MORALES P, TAPIA E, PEDRAZA-CHAVERRI J.β-hydroxybutyrate:a signaling metabolite in starvation response?[J].Cellular Signalling, 2016, 28(8):917-923.

[15] PAOLI A, BOSCO G, CAMPORESI E M, et al.Ketosis, ketogenic diet and food intake control:a complex relationship[J].Frontiers in Psychology, 2015, 6:27.
[16] KADOWAKI T, YAMAUCHI T, KUBOTA N.The physiological and pathophysiological role of adiponectin and adiponectin receptors in the peripheral tissues and CNS[J].FEBS Letters, 2008, 582(1):74-80.

[17] WANDERS D, GRAFF E C, WHITE B D, et al.Niacin increases adiponectin and decreases adipose tissue inflammation in high fat diet-fed mice[J].PLoS One, 2013, 8(8):e71285.
[18] BEHRE C J.Adiponectin:saving the starved and the overfed[J].Medical Hypotheses, 2007, 69(6):1290-1292.

[19] OKAMOTO Y, KIHARA S, FUNAHASHI T, et al.Adiponectin:a key adipocytokine in metabolic syndrome[J].Clinical Science, 2006, 110(3):267-278.

[20] 邹华围.饥饿后恢复饲喂和β-羟基丁酸灌注对牦牛脂肪代谢的影响[D].博士学位论文.雅安:四川农业大学, 2019. ZOU H W.Effects of refeeding and β-hydroxybutyrate infusion on lipid metabolism in starved yak[D].Ph.D.Thesis.Ya'an:Sichuan Agricultural University, 2019.(in Chinese)
[21] 郭红芳, 昝林森, 孙永刚.牛前体脂肪细胞的分离培养及诱导分化[J].西北农林科技大学学报(自然科学版), 2014, 42(2):1-6, 12. GUO H F, ZAN L S, SUN Y G.Primary culture and differentiation of bovine preadipocytes[J].Journal of Northwest A & F University (Natural Science Edition), 2014, 42(2):1-6, 12.(in Chinese)
[22] 陈涛, 马立霞, 李志鑫, 等.新西兰兔肌内和皮下前体脂肪细胞原代培养与诱导分化的研究[J].中国畜牧杂志, 2019, 55(6):39-44. CHEN T, MA L X, LI Z X, et al.Comparative study on primary culture and induced differentiation of intramuscular and subcutaneous preadipocytes of New Zealand rabbits[J].Chinese Journal of Animal Science, 2019, 55(6):39-44.(in Chinese)
[23] 赵蕾, 郑美丽, 杨梅, 等.3T3-L1前体脂肪细胞诱导分化方法优化的初步探讨[J].首都医科大学学报, 2019, 40(2):226-231. ZHAO L, ZHENG M L, YANG M, et al.Optimization of induction and differentiation of 3T3-L1 preadipocyte[J].Journal of Capital Medical University, 2019, 40(2):226-231.(in Chinese)
[24] NISHITANI S, FUKUHARA A, SHIN J, et al.Metabolomic and microarray analyses of adipose tissue of dapagliflozin-treated mice, and effects of 3-hydroxybutyrate on induction of adiponectin in adipocytes[J].Scientific Reports, 2018, 8(1):8805.
[25] CARLSON L A.Studies on the effect of nicotinic acid on catecholamine stimulated lipolysis in adipose tissue
in vitro[J].Acta Medica Scandinavica, 1963, 173:719-722.
[26] DE PERGOLA G, CIGNARELLI M, CORSO M, et al.
D-(-)-beta-hydroxybutyrate inhibits catecholamine-stimulated lipolysis and decreases beta-adrenoceptors' affinity in human fat cells but not in lymphomonocytes[J].Acta Endocrinologica, 1990, 122(4):450-454.

[27] TAGGART A K P, KERO J, GAN X D, et al.(
D)-beta-hydroxybutyrate inhibits adipocyte lipolysis via the nicotinic acid receptor PUMA-G[J].Journal of Biological Chemistry, 2005, 280(29):26649-26652.

[28] DE OLIVEIRA CAMINHOTTO R, KOMINO A C M, DE FATIMA SILVA F, et al.Oral β-hydroxybutyrate increases ketonemia, decreases visceral adipocyte volume and improves serum lipid profile in Wistar rats[J].Nutrition & Metabolism, 2017, 14:31.
[29] KENÉZ A, LOCHER L, REHAGE J, et al.Agonists of the G protein-coupled receptor 109A-mediated pathway promote antilipolysis by reducing serine residue 563 phosphorylation of hormone-sensitive lipase in bovine adipose tissue explants[J].Journal of Dairy Science, 2014, 97(6):3626-3634.

[30] DENG Q H, LIU G W, LIU L, et al.BHBA influences bovine hepatic lipid metabolism via AMPK signaling pathway[J].Journal of Cellular Biochemistry, 2015, 116(6):1070-1079.

[31] ZHANG M, ZHANG S Q, HUI Q, et al.β-hydroxybutyrate facilitates fatty acids synthesis mediated by sterol regulatory element-binding protein1 in bovine mammary epithelial cells[J].Cellular Physiology and Biochemistry, 2015, 37(6):2115-2124.

[32] FAJAS L, SCHOONJANS K, GELMAN L, et al.Regulation of peroxisome proliferator-activated receptor gamma expression by adipocyte differentiation and determination factor 1/sterol regulatory element binding protein 1:implications for adipocyte differentiation and metabolism[J].Molecular and Cellular Biology, 1999, 19(8):5495-5503.

[33] TRUJILLO M E, SCHERER P E.Adiponectin-journey from an adipocyte secretory protein to biomarker of the metabolic syndrome[J].Journal of Internal Medicine, 2005, 257(2):167-175.

[34] PLAISANCE E P, LUKASOVA M, OFFERMANNS S, et al.Niacin stimulates adiponectin secretion through the GPR109A receptor[J].American Journal of Physiology:Endocrinology and Metabolism, 2009, 296(3):E549-E558.
[35] 尉小强.饥饿对牦牛糖异生和脂肪代谢的影响研究[D].硕士学位论文.雅安:四川农业大学, 2015. WEI X Q.Effects of starvation on gluconeogenesis and fat metabolism of yak[D].Master's Thesis.Ya'an:Sichuan Agricultural University, 2015.(in Chinese)